Campbell, . — Studies on some Javanese Anthocerotaceae. II. ioi 
As we have pointed out, this formation of the sporogenous tissue from 
the central part of the sporogonium has not been confirmed by the 
writer’s study of N. javanicus , although the slight development of the 
columella in some of the small sporogonia indicates the possibility of a 
complete suppression of the columella in the poorly nourished sporogonia. 
A comparison of the sporogonium of No to thy l as is perhaps best 
made with Sphaerocarpas or even better with Cyatkodium. 1 The latter 
is especially interesting, as the foot of the sporogonium, unlike that 
of any other liverwort, produces root-like extensions like those of the 
typical Anthocerotaceae. It is also worthy of note that the Targionieae, 
and especially Cyatkodium , are characterized by remarkably large chro- 
matophores, those in C. foetidissimmn being sometimes reduced to four 
in some of the assimilative cells. The writer has examined this point 
in specimens collected in Java, and the resemblance to the chromatophores 
of Megaceros is sufficiently striking. Should these resemblances in 
Cyatkodium and the Anthocerotaceae prove to be anything more than 
coincidences it would show that the affinities of the Anthocerotaceae 
are rather with the Marchantiales than with the Jungermaniales. The 
relationship in either case is certainly remote, and for the present at 
least it will probably be best to regard the Anthocerotaceae as sufficiently 
distinct from the true Hepaticae to form a special class, Anthocerotes, 
as was suggested by Howe . 2 
EXPLANATION OF PLATES IX AND X. 
Illustrating Professor Campbell’s Studies on Anthocerotaceae, II. 
PLATE IX. 
Fig. l. Dendroceros sp. { A). Thallus. x 3. 
Fig. 2. Apex of a shoot with antheridial branch, oA x 3. sp , young sporophytes. 
Fig* 3* two ripe sporogonia of the same, x 3. col , columella ; in, involucre. 
Fig. 4. Plant of Dendroceros sp. (B) with two sporogonia, sp. x 3. 
Fig. 5. A ripe sporogonium of the same, x 3. 
Fig. 6. Cells of the wings of the thallus of B, showing the large lacunae, x no. 
Fig. 7. A single cell showing the chromatophore with the included pyrenoid, p. x 280. 
n, nucleus. 
Fig. 8. Apical region of the thallus of B, longitudinal section, x 280. 
Fig. 9. A similar section of A ; ?, archegonia ; l, mucilage cavities. 
Fig. 10. Young archegonium of A. x 480. 
Fig. 11. Young archegonium of B. x 480. 
Fig. 12. Open archegonium of B. x 280. 
1 Lang, On the Morphology of Cyatkodium , Ann. of Bot., xix, p. 41 1, 1905. 
3 Mem. Torrey Botanical Club, vii, 1899. 
