326 
ME. ST. G-EOBGrE MIYAET ON THE SKELETON OE THE PEIMATES. 
metacarpal. It almost equals it in length in Lagothrix, Cebus, Pithecia, Chryso 
thrix, Lemur, Arctocebus, and sometimes in Hapale *. It slightly exceeds it in Bra- 
chyurus, Nyctipithecus, Galago, Loris, and Perodicticus, and very largely so in Tarsius 
and Cheiromys. 
Its proportion to the manus is greatest in the two last-mentioned genera (about as 
34 to 100), in all the rest it is more than a quarter, except in Ateles, Cynocephalus, 
Cercopithecus, Troglodytes, Man, and last of all, Arctocebus, where it is as 21 "5 to 100. 
It is at its greatest absolute length in Simia. 
The second phalanx is at its greatest absolute length in Simia and the Chimpanzee. 
It is always more than half the length of the proximal phalanx, except in Perodic- 
ticus and Cheiromys, where it is a little less. It is never, however, nearly so long as 
the first phalanx. 
The third phalanx of the third digit is, like the other phalanges, absolutely longest 
in Simia. 
It is always shorter than the second phalanx, and is less than half its length in the 
Chimpanzee, Orang, Hylobates, Semnopithecinse, Macacus, Pithecia, Nyctipithecus, 
Chrysothrix, Indris, Lemur, 'Loris, and Cheiromys. In Tarsius alone is it less' than one- 
third of the length of the second phalanx. 
The phalanges always shorten successively, except that the second phalanx of the 
fourth digit is longer than the proximal phalanx of the index in Galago and the Nyc- 
ticebinse (especially, of course, Perodicticus and Arctocebus), and that it is longer than 
the proximal phalanx of the fifth digit also in Cheiromys. 
Digits without their Metacarpals. 
Th e pollex thus measured is absolutely longest in Man when of average size. 
As compared with the whole length of the manus, it is greatest in Arctocebus (about as 
35 to 100), then in Hapale, Chrysothrix, and Man. In all it is more than one-fifth the 
length of the manus, except in the Simiinse and Semnopithecinse, Nyctipithecus, and 
Galago. It is less than a twentieth in Colobus f . 
The pollex is never the longest digit of the manus, but, except in Perodicticus and 
Arctocebus, it is always the shortest one. 
The index is never the longest digit of the manus, but in the two last-mentioned 
genera it is the shortest one. It is decidedly the shortest, except the pollex in the Le- 
muridse and Cheiromys. Tt projects furthest distad in none ; it does so least of the 
* According to Dr. Lucae, the first phalanx is equal to the metacarpal in length in Hapale and Galago 
( loc . cit. p. 320). 
t Dr. Lucae, loc. cit. p. 318, says that the pollex without its metacarpal does not by a good deal attain in the 
tailed Apes the proportionate length which it reaches in the Simiinse. I have found all the lower Simiidae to 
exceed all the Simiinse in this respect except T. niger, and Dr. Lucae, in his Table B., gives for the proportion 
to the manus at 100, 22 & 24 to Cynocephalus, while to H. Leuciseus (which has the highest proportion of his 
Simiinas) he only assigns 21-7 (see pp. 307 &317). 
