356 
MR. ST. GEORGE MIVART ON THE SKELETON OE THE PRIMATES. 
cuneiforme, and thus stamped with an oblique depression, or, if turned still further, 
with a transverse one. 
The shaft of the bone is generally nearly straight, but in the Simiinse it has a twist, 
which seems to disappear, or almost so, with the acquisition of a nearly transversely ex- 
tended concavity in the articular surface. Thus in the Simiinas the distal end of the bone 
can assume a position which would be impossible without this twist. 
This distal end of the hone is rarely so large as the proximal one, and never predomi- 
nates over it, as is the case with the homotypal parts in the pollex. The angle formed 
by an axis piercing it from side to side transversely, with another similarly traversing 
the heads of the other metatarsals *, is very rarely (only in Man) so extremely obtuse as 
to approach 180°. 
In the Simiinse, as well as in the lower forms, this angle is much more acute, approxi- 
mating to 90°. 
This metatarsal is the longest one of the whole pes in Galago and Arctocebus. It is 
shorter than the other metatarsals in all the other forms except Loris, Perodicticus, and 
Tarsius ; though in Indris it is very little so. 
Second Metatarsal . — This is sometimes the absolutely longest metatarsal of all, as 
has been said, but in a certain form (Arctocebus) it is the absolutely shortest of 
the order. 
It is the longest of all in the same pes in Man, and (if we exclude the backwardly 
extending process of the base of the fifth metatarsal) also in the Simiinse and Lemur, 
and at least sometimes in Ateles and Lagothrix. 
It is the shortest metatarsal in Arctocebus, Perodicticus, and Tarsius ; and it is the 
shortest except that of the hallux in the Semnopithecinae, Pithecia, Nyctipithecinee, Ha- 
pale, Indris, and Cheiromys. 
It projects more forwards (i. e. distad) than the three metatarsals external to it in 
Man, the Simiinae, Ateles, and Lagothrix. 
It projects less than those do in Pithecia, Chrysothrix, Hapale, Indris, Tarsius, and 
Cheiromys. 
It is longer than the second metacarpal in all except the Simiinae, Tarsius, and 
Cheiromys. 
The proximal end has an articular surface, which is flat, as in Man, or concavo-con- 
vex, as in the other Anthropoidea, or convex only, as in the Lemuroidea. 
Sometimes (rarely), as e. g. in Lemur and Loris, it meets the proximal end of the 
fourth metatarsal beneath the ecto-cuneiforme. 
The proximal end extends further back than the base of the third metatarsal. In Man, 
Troglodytes (only slightly so in the Gorilla), Hylobates, the lower Simiidse, sometimes in 
Cebus, in Pithecia, Nyctipithecus, Hapale, in the Lemuroidea generally, and greatly 
so in Lemur. 
Third Metatarsal . — This is absolutely longest in Simia. 
* The plantar angulation of Professor Huxley. 
