ME. ST. GEOEGE MIVAET ON THE SKELETON OE THE PEIMATES. 
357 
It is the longest of all in the same pes in the lower Simiidse, sometimes in Cebns, in 
Mycetes, Perodicticus, and Cheiromys. 
It is never the shortest of the pes, but it is longer than the third metacarpal in all 
except the Simiinse and Cheiromys, though it is almost the same length in Galago and 
Tarsius. 
It projects most forwards, of the four outer metatarsals, in the lower Simiidee, Cebus, 
Mycetes, the Nyctipithecinse, and Lemuroidea. 
The proximal articular surface is flat in Man, and more or less convex in the other 
forms. 
Fourth Metatarsal . — This is the longest one of the pes in Pithecia, and the Nycti- 
pithecinae, Hapale, Indris, and Tarsius. 
It is the shortest metatarsal in Loris and Galago. 
It projects the most forwards of all the metatarsals in Pithecia and Hapale. 
It is longer than the fourth metacarpal in all except the Simiinse ; but the length of 
these two bones is nearly the same in Galago and Arctocebus. 
The proximal end is almost always strongly convex, except in Man. 
Fifth Metatarsal . — When the backwardly-projecting process of its base is included, 
this metatarsal is the longest of the pes in the Gorilla, many Cebidse, Cheiromys, and 
sometimes in Man. 
Without including that process, it is the shortest of the four outer metatarsals in 
Man, the Simiinse, Ateles, Lagothrix, Lemur, and Loris. 
Except that of the hallux, it projects less forwards than any other of the metatarsals 
in Man, the Simiidse, the Cebinse, Mycetes, Nyctipithecus, Lemur, Galago, and Loris. 
It is longer than the fifth metacarpal in all except the Chimpanzee, Simia, and Hylo- 
bates ; but it is very little so in Arctocebus and Perodicticus, while in Brachyurus, Nyc- 
tipithecus, Chrysothrix, and Hapale it is more than double the length of the fifth meta- 
carpal. 
Its proximal end is in general strongly convex. A process projects backwards from 
the outside of the proximal end of the fifth metatarsal. This is at its maximum in Man 
and the Gorilla. It is smaller relatively, as well as absolutely, in most others, espe- 
cially in the Lemuroidea. 
Phalanges. 
The hallux has always two phalanges, except, as is well known, in Simia, where there 
is often but one. 
Each of the other digits has three distinct phalanges, except in Man, where generally 
the ultimate and penultimate phalanges of the fifth digit become anchylosed together. 
The proximal phalanx of the hallux is absolutely greatest in Man, but the Gorilla 
approaches him very closely * in this respect. 
Professor Owen has found it to equal that of Man (Comp. Anat. of Yertebrates, vol. ii. p. 551). 
3 c 2 
