ME. ST. GEOEGE MIVABT ON THE SKELETON OE THE PEIMATES. 
361 
small in Indris, Arctocebus, and Loris. In most, however, it is smaller than in Man ; but he 
is exceeded in this respect by Mycetes, Tarsius, Chrysothrix, Cheiromys, Nyctipithecus, 
and Hapale, in which the difference between the proportions becomes successively 
greater, the excess of the longest digit of the manus over the longest metacarpal being 
in the last-mentioned genus more than four-fifths greater than that of the longest digit 
of the pes over the longest metatarsal. 
Digits with their Metatarsals. 
The hallux thus estimated is absolutely longest in Man. 
Its proportion to the spine is far greatest in Tarsius (more than a quarter), and then 
in Cheiromys and Hylobates, Indris, and Ateles, in all of which it is more than one-fifth 
the length of the spinal column. In Colobus and Hapale, on the other hand, it is but 
little more than one-tenth. 
Its length, compared with that of the entire pes, is greatest in Arctocebus, Loris, and 
Indris, where it is more than one-half the length of the latter — a proportion it nearly 
attains in Man, and sometimes in Hylobates, while the Chimpanzee follows closely. In 
all the rest it is more than 33 to 100, except in Hapale and the Semnopithecinae, where 
it is a little less, and Simia, where it is scarcely more than a quarter. 
The hallux, when brought beside the index digit, attains to its extremity in Arcto- 
cebus ; sometimes beyond its extremity in Man : to the middle of the distal phalanx, 
or rather beyond it, in the other Nycticebinse, Galago, Tarsius, and sometimes in Man : 
to the middle or near the distal end of the second phaJanx of the index in Lemur and 
Indris : to the proximal end of the second phalanx in the Chimpanzee and Cheiromys : 
to the distal end of the proximal phalanx in the Gorilla, sometimes Hylobates, 
Cynocephalus, Pithecia, and Nyctipithecus : to the middle, or nearly so, of the proximal 
phalanx in Hylobates (sometimes), the Cebinae, Mycetes, Chrysothrix, and some lower 
Simiidae : to a little beyond the base of the proximal phalanx in the Semnopithecinae 
and Hapale : not nearly to the distal end of the metatarsal of the index in Simia. 
The extent to which the hallux extends with regard to the index of the pes, when 
compared with the extent to which the pollex projects forwards beside the index of the 
manus in the same individual, is as follows ; — 
Almost always the hallux projects further than the pollex (omitting Arctocebus and 
Perodicticus). 
The reverse condition, however, obtains largely in Hapale, in a less degree in some 
Cebidae, e. g. Lagothrix, Mycetes, and also in Simia. 
In most of the Cebidae the relative extension is about equal ; but in Man and Tarsius 
Museum had been, wrongly articulated ; but in Yan Campen’s memoir, before referred to, bis plate represents 
the longest metacarpal as about equal to the longest metatarsal in length, while the longest digit of the pes 
decidedly exceeds that of the manus. 
