Bower . — Studies in the Phytogeny of the Filicales . VI. 5 
Venatio Anaxeti. The fertile leaves have narrower lobes, but still with the 
well-marked midrib. Their lower surfaces are covered right and left of the 
midrib by a dense and continuous sorus of Acrostichoid character. The 
drawings published by Sir William Hooker 1 adequately represent the main 
characters; but in order to show the chief habit-features of the plant, 
a photograph has been made of the finest specimen sent from Darjeeling, 
the plant being about three feet in height (Plate I, Fig. 2). 
Anatomy. 
The adult leaf-trace of Leptochilus tricuspis consists of five, or some- 
times of six, strands, arranged in a horseshoe, with the open side facing 
directly towards the apex of the axis. The two marginal strands are 
Text-fig. i. Series of transverse sections of the rhizome of Leptochilus tricuspis , including 
a leaf-insertion, x 4. 
larger than the rest, and show indications of the usual recurved hooks. 
The vascular system of the axis is in the internodes disposed in a ring 
of strands of unequal size, circular or slightly oval in outline, with occa- 
sional fusions, and here and there insertions of root-strands directed laterally 
or downwards (Text-fig. 1, a, b). When a leaf-trace is inserted, the distance 
between the median strands on the upper side increases, and these strands, 
which are usually of large size, rotate on their axes, so that the edges of their 
xylem-plates are directed towards the incoming trace (c, d). Upon these 
the marginal strands of the leaf-trace insert themselves (d, e, f), while 
the smaller median strands fusing irregularly (e,f) enter the vascular ring. 
The whole construction is such as would naturally follow if a solenostelic 
axis and a united leaf-trace, such as. is seen in Dipteris or Metaxya , were 
1 Sp. Fil., vol. v, Tab. CCCIV. 
