On the Occurrence of Intrafascicular Cambium in 
Monocotyledons. 
BY 
. AGNES ARBER. 
With three Figures in the Text. 
Introduction. 
A N impression seems to prevail among botanists that the occurrence 
of intrafascicular cambium in Monocotyledons is extremely rare, 
except perhaps in seedlings. It may therefore be worth while to collect 
together some of the scattered observations on this subject, which tend to 
show that an ephemeral cambium is, in reality, a widespread anatomical 
feature of this group, and also to record a few fresh examples and to figure 
and describe briefly a case in which the cambium can be particularly well 
observed. 
I am indebted to Miss Saunders, Director of the Balfour Laboratory, 
for facilities for anatomical work, and to the Newnham College Fellowship 
Committee for a grant towards the expenses of this and other researches. 
I wish also to express my thanks to Mr, Boodle, for his help in connexion 
with the literature, and to Mr. Lynch, of the Cambridge Botanic Garden, 
for his kindness in supplying me with material. 
Review of the Literature. 
The earliest observation on the occurrence of intrafascicular cambium 
in Monocotyledons appears to be that of Russow, 1 who, in 1875, described, 
in the vascular bundle of Hemerocallis fulva , ‘ eine Andeutung eines Cam- 
biumstreifens \ This was followed in 1884 by Godfrin’s 2 record that in the 
veins of the cotyledon of Latania borbonica (Palmae) an active cambial zone 
occurred. Two years later Mobius 3 observed traces of a cambium in the 
stems of Lister a ovata and Orchis maculata , and found that Limodorum 
abortivum was distinguished by somewhat more prolonged cambial activity. 
In 1888 the subject was placed on a much broader basis by Froken 
Sigrid Andersson’s 4 researches, which, owing to their publication in 
1 Russow, F. (1875). 2 Godfrin, J. (1884), p. 47 and PI. Ill, Fig. 43. 
3 Mobius, M. (1886). 4 Andersson, S. (1888). 
[Annals of Botany, Vol. XXXI. No. CXXI. January, 1917.] 
