44 
Arber . — On the Occurrence of 
another inflorescence axis, gathered on April 27, as many as ten undif- 
ferentiated elements have been traced in a single radial row. In Fig. 2 
a bundle is represented from a lower level in the axis gathered on May 7. 
Cambial activity is still in progress, but part of the secondary phloem 
has become modified into large sieve-tubes and small companion cells, thus 
somewhat obliterating the radial arrangement. Fig. 3 shows a bundle 
from the flowering region of an older inflorescence, gathered on May 28. 
Here cambial activity seems to have completely, or almost completely 
ceased. In these figures it will be observed that little or no secondary 
xylem is formed ; the cambium seems to confine its activities, mainly 
if not entirely, to the production of secondary phloem. The same pecu- 
liarity has already been recorded for other members of the Liliaceae by 
Andersson. 
Meristematic activity is not rigidly restricted to a single layer of 
initial cells. Dividing nuclei seem most frequently to occur near the 
innermost or xylem end of the radial files ; they have been noticed in 
different cases in the first, second, third, fourth and sixth cells, counting 
from the inner end. In one case nuclei were seen dividing simultaneously 
in the fourth and sixth cells of the same file, and, in another case, in the first 
and third. 
In addition to Eremurus , a few other genera among the Liliaceae have 
been examined. In the case of shoots of Asparagus officinalis , L., at the 
stage at which they are gathered for market, an ephemeral cambium 
was detected in the apical region. In such a shoot, gathered on May 1, 
transverse sections across the * head ’, at a distance of 1 cm. from the apex, 
showed the metaphloem to consist of radial rows of undifferentiated cells, 
uniformly filled with cytoplasm. Traces of the same arrangement were 
visible at the base of the * head \ but, 2*5 cm. lower, the differentiation into 
sieve-tubes and companion cells had completely masked all indications of 
radial grouping. 
Distinct traces of cambium have also been observed in a young inflores- 
cence axis of Nothoscordmn fragrans , Kunth, gathered on May 18. The 
bundles are more or less Y-shaped, and the cambium follows a curved line. 
In the young inflorescence axis of Hemerocallis fulva , L., cambium has 
also been recognized. Russow, 1 as has already been mentioned, noted more 
than forty years ago the occurrence of cambium in another species, H . 
Jlava , L., so the present observation merely confirms its existence in 
this genus. 
Summary. 
In this paper the literature on intrafascicular cambium in Monocotyle- 
dons is briefly reviewed, and it is recorded that, in addition to the cases 
1 Russow, E. (1875). 
