the Mycorrhiza of the Marattiaceae. 95 
separated localities as Eastern Asia (Java, South-West China), Australasia, 
and Ceylon. 
Diagnosis : 1 
Stigeosporium , gen. nov. 
Mycelium ramosum ex hyphis inter- et intracellularibus continuis 
rarissime septatis constans ; haustoriis numerosis, extremitatibus in ramulis 
radiatis valde dissectis; sporis perdurantibus solitariis plerumqueglobosis raro 
subglobosis et cet., membrana crassissima irregulariter intenseque colorata. 
Differt a Phytophthora , cui arete affine, habitu symbiotico, qua de causa 
nulla conidia normalia producuntur. 
Sp. unica. 
Stigeosporium M arattiacearum^ West, sp. nov. 
Hyphae primum hyalinae demum flavo-brunneae vacuatae 1-12 [x 
crassae ; sporis vel intercalaribus vel terminalibus plerumque globosis 
33-45 ix diametro, raro subglobosis vel ovoideis vel pyriformibus ; exosporio 
tenuissimo hyalino levi, mesosporio crassissimo 6 fx crassitudine minute 
punctulato flavo irregulariter intenseque colorato, endosporio tenue 3*5 ix 
crassitudine. 
Hab. In radicibus subterraneis quae ad quasdam marattiacearum 
generum orientalium omnium adhuc cognitorum species pertinent. 
Distrib. Asia orientalis, Australasia, Zeylania. 
B. On the Endophyte of the Roots of Danaea alata, Sm., 
and of Danaea nodosa, Sm. 
The principal roots of both species of Danaea examined by the present 
writer are characterized by the presence of a zone of fibrous cells which 
is separated from the epidermis by a few layers (2-6) of large parenchyma- 
tous cells which constitute the outer cortex (Text-fig. 9 ; PI. Ill, Figs. 3 and 
4) : in the smaller roots, however, no fibrous zone is found. 
In other respects the roots of Danaea very closely resemble those 
of the other genera of Marattiaceae. 
An endotrophic fungus was found in nearly every root examined, 
irrespective of age and size ; it is invariably present in the primary and first 
few adventitious roots of the young sporophyte, but a few of the older roots 
were found to be free from fungal mycelium. 2 
The mycelium is irregularly distributed throughout the cortex of the 
smaller roots which do not possess a cortical zone of stereome, but in the 
larger roots the mycelium is always confined to the few layers of parenchyma 
outside the fibrous zone of the cortex (Text-fig. 9 ; PI. Ill, Figs. 3 and 4). 
1 The above diagnosis is an extended form of that which the author published as a note in 
Annals of Botany, vol. xxx, 1916. 
2 An endophyte exhibiting similar characters was observed in the cells of the prothallia. 
