96 
West. — On Stigeosporium Marattiacearum and 
In Danaea also, the presence of the endophyte does not in any way 
alter the external appearance of the roots. 
Infecting hyphae enter the root either through ordinary epidermal 
cells or through the root-hairs (Text-fig. 9). 
The hyaline non-septate copiously branched mycelium consists for the 
most part of fine intracellular hyphae which are fairly uniform in diameter 
(2-4 m)* These hyphae have a delicate membrane and finely granular con- 
tents in which no nuclei can be identified with certainty (Text-fig. 9 ; PL III, 
Fig. 4, myc . ; and cf. Stigeosporium). 
Text-fig. 9. Part of a transverse section of a moderately large root of Danaea alata , Sm., 
showing endophyte {end.) in outer cortical cells, fibr. 1 . = cortical zone of stereome ; n. — nucleus 
of host-cell ; r.h. = root-hair ; s.gr. = starch grains ; sp. = foreign spore-like bodies ; v. — vesicle ; 
x. = disorganized 1 arbuscules ’. x 300. 
From the relatively few intercellular hyphae ‘ arbuscules 5 essentially 
similar to those already described for Stigeosporium Marattiacearum are 
produced. These rapidly collapse, their disorganized remains forming 
irregular structureless granular masses within the host-cell (Text-fig. 9 
PL III, Figs. 3 and 4). Vesicles are occasionally found : they are generally 
globular in form and are never delimited by a septum from the supporting 
hypha, upon which they occupy a terminal position (Text-fig. q,v ; PI. Ill, 
Fig. 4, v). Intercalary vesicles were never observed in this endophyte. The 
hyphae often form peculiar coiled bunches within the mycorrhizal cells 
(Text-fig. 9) : the real significance of this phenomenon, which is not 
uncommon among endotrophic mycorrhizal fungi, is not yet understood. 
So far as it is possible to judge from an examination of sections of 
infected roots, the hyphae of this endophyte exert no deleterious effect upon 
