the Mycorrhiza of the Marattiaceae. 97 
the invaded cells, the only obvious result of their presence being the 
complete disappearance of starch from such cells. 
Unfortunately, no reproductive bodies were observed, consequently 
the systematic position and relationships of this endophyte could not, even 
approximately, be determined. No living material of either species of 
Danaea was available for investigation ; it was therefore impossible to 
carry out cultural experiments with this fungus. 
That the endophyte which is associated with the roots of Danaea spp. 
is quite distinct from Stigeosporinm is proved by the following characters, 
which belong only to the former : 
1. Mycelial characters . 
A. Uniform diameter of the hyphae; 
B. Comparative rarity of intercellular hyphae ; 
C. The hyphal wall does not materially thicken with age ; 
D. Absence of distinct nuclei from contents of the mycelium. 
2. Apparent absence of definite reproductive bodies . 
3. Absence of vesicles of irregidar form. 
Summary. 
1. A new fungus, Stigeosporinm Marattiacearum , which forms an 
endotrophic mycorrhiza with roots of certain genera (Angiopteris, Archangio- 
pteris , Kaulfussia , Marattia) of Marattiaceae, is described, and its biology, 
probable life-history, and systematic position discussed. 
2. An unnamed mycorrhizal fungus, which enters into association with 
roots of the genus Datiaea , is briefly described. 
I take this opportunity of expressing my sincere thanks to Professor 
J. B. Farmer, F.R.S., for many helpful suggestions, and for allowing me to 
carry out the present investigation in his laboratory. 
Royal College of Science, 
South Kensington, S.W. 
Literature. 
1 . Bernard, Noel : Sur quelques germinations difficiles. Rev. gen. de Bot., t. xii, 1900. 
2 . : Etudes sur la tuberisation. Rev. gen. de Bot., t. xiv, 1902. 
3 . : La germination des Orchidees. C. R. Acad. Sci., t. cxxxvii, 1903. 
4 . — : Recherches experimentales sur les Orchidees. Rev. gen. de Bot.,t. xvi, 1904. 
5 . — : Le champignon endophyte des Orchidees. C. R. Acad. Sci., t. cxxxviii, 
I9 ° 4 * 
6. - — ■: Nouvelles especes d’endophytes d’Orchidees. C. R. Acad. Sci., t. cxl, 1905. 
7 . : Symbioses d’Orchidees et de divers champignons endophytes. C. R. Acad. 
Sci., t. cxlii, 1906. 
■: L’evolution dans la symbiose des Orchidees et leurs champignons com- 
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