156 Blackburji. — On the Vascular Anatomy of 
strongly that of a monocotyledon. The seedlings of the genus have been 
described by M. Mansion in the paper referred to above. The young plants 
differ from those of Aquilegia in having petioles to the plumular leaflets. 
The anatomical features of the four species examined, namely, T. javanicum, 
T. flavum , T. glaucum , and T. adian tifolium , are very similar in the seedling 
stage. The vascular cylinder consists of a ring of bundles, and at each node 
three separate leaf-trace strands are inserted. At the cotyledonary node 
are the usual six plumular bundles, besides the entering cotyledonary traces, 
which, though single higher up, show their double nature at this level. The 
anatomy thus closely resembles that of the genus Aquilegia, and the general 
type described for the order. 
Anemone. The adult anatomy in this genus has been studied in detail 
by M. Janczewski 1 and less fully by M. Marid. The latter, in summing 
up for the genus, says that the cauline bundles are in one circle though of 
different sizes . 2 
These strands may be surrounded by individual endodermal sheaths. 
Medullary bundles are found in some species in addition to the normal 
ring. 
I understand M. Janczewski to state that the habit of the stem has 
definite anatomical consequences influencing the presence or absence of 
endodermis, cambium, and mechanical tissue. Mr. Sinnott speaks of the 
genus as being characterized by a multilacunar leaf-trace insertion . 3 
In the young hypogeal stem of A. pennsylvanica, Professor Jeffrey has 
discovered a stelar tube with internal and external phloeoterma communi- 
cating through the leaf-gaps,. 
In all but two of the species examined the seedling anatomy corre- 
sponds very closely to that of Aquilegia , though the morphological features 
show a large range of variation. Most species of Anemone have a cotyle- 
donary tube, and associated with this, in A. apennina, there is a tuberous 
hypocotyl. In another species, A. nemorosa, the tuberous habit is found in 
connexion with the hypogeal germination. 
Anemone montana . This is a small seedling in general habit resembling 
Aquilegia , but with trifid foliage leaves. The internal structure is corre- 
spondingly similar to that of Aquilegia . Cambial growth is not present in 
the strands at the leaf base, but is clearly marked and quite diagrammatic 
in the axis at an early stage. 
The lateral strands of the trifascicular trace are nearer the median one 
at the node than in the last two genera, so that those of subsequent leaves 
do not so frequently overlap. 
1 Janczewski: Etude morphologique sur le genre Anemone . Rev. Gen. de Bot., vol. x 
1898. 
2 Loc. cit., p. 167. 
3 Sinnott, E. W. : The Anatomy of the Node as an Aid in the Classification of Angiosperms. 
Am. Journ. of Bot., vol. i, July, 1914, p. 312. 
