174 Blackburn . — On the Vascular Anatomy of 
plant. The seedling shows only the usual three. The epicotyledonary 
anatomy is almost identical with that of Anemone. Just above the cotyle- 
donary node the plumular bundles are as usual six in number. At the 
node the plumular tissue divides in the intercotyledonary plane, but leaves 
in the middle the protoxylem elements of the median strands. The double 
bundles of the cotyledons orientate themselves in the usual manner to give 
a diarch arrangement, and the plumular protoxylem groups thus lie on the 
same radius as the two phloem strands. Both pith and plumular strands 
die out at a somewhat lower level, and thus root structure is arrived at. 
B. heteropoda is similar to B. aquijiornm. 
B. lyceum and B. aristata show similar epicotyledonary features. In 
the hypocotyl the four cotyledonary phloem groups persist in the diagonal 
planes, and the protoxylem groups from the median strands of the first two 
plumular leaves penetrate farther into the hypocotyl than in the other forms. 
Lardizabalaceae. 
Only Decaisnea Fargesii has been obtained of this order. This and the 
forms to be described in the following orders are trees, and the seedlings 
show certain differences from the herbaceous forms of Ranunculaceae. 
The seedlings are larger and the internodes are elongated. The leaf-trace 
insertion is as usual trilacunar, but there is a larger number of strands in 
the vascular ring. 
In Decaisnea the seedling is large with long-stalked, irregularly oval 
cotyledons. The first foliage leaf is compound, with three leaflets, and the 
second has five. The bases of the foliage leaves are not sheathing, as they 
are in Ranunculaceae. 
The plumular internodes are long, while the phyllotaxis is the usual 
two-fifths. Axillary buds are present both in connexion with the cotyle- 
dons and foliage leaves. 
The vascular cylinder of the axis consists of a ring of about twenty 
bundles which are connected at an early age by cambium. The leaf trace 
has the usual trilacunar insertion, but the median strand is made up of three 
at its point of entry into the ring. While still in the cortex the leaf-trace 
strands divide up, the lateral ones each into two and the central one form- 
ing a ring of about seven bundles. These latter separate into two adaxial 
and five abaxial strands, and rearrange themselves so that in the base of the 
petiole there is a complete cylinder of bundles. At the cotyledonary node 
the plumular strands are reduced to two flattened arcs of five bundles on 
either side of the cotyledonary plane. The two separate bundles of the 
cotyledons, derived by fusion of four at a slightly higher level, insert them- 
selves at the ends of these arcs. The ten plumular bundles reduce to six, 
and the arcs close up to form a ring which persists far down the hypocotyl 
and is similar to that found in the epicotyledonary region. Diarch root 
