West. — A Contribution to the Study of the Marattiaceae. 377 
vegetatively produced plants tend in their development to pass through 
stages in elaboration similar to young plants which take their origin from 
a zygote (cf. Jones, 37 , p. 27). This statement also holds good for the 
adventitiously produced plants of Danaea nodosa. At the apex of the 
stem of the adventitious bud there is a single elongated initial cell, similar 
in every respect to that found at the stem-apex of the sporeling (cf. Text- 
fig. io, B, with Text-figs. 20, B, and 21, B). Moreover, the second leaf of the 
adventitious bud is formed almost exactly opposite to the first leaf (cf. Text- 
fig. io, A, with Text-fig. 3, B). Not only does the adventitious bud actually 
arise upon the basal pulvinus of the petiole (although very close to the stem), 
but its vascular supply is derived directly from one of the foliar traces (Text- 
fig. 8 , b) ; it cannot therefore be regarded as a branch of the parent stem. 
Text-fig. io. a. Median longitudinal section through the apex of an adventitious bud 
of Danaea nodosa , Sm. a., apical cell of the stem ; A/. 1 , l.tr, traces of the first and second leaves 
respectively, x 8. b. Apical cell of same more highly magnified, x 360. 
Vascular Anatomy. In the vegetatively produced plant the vascular 
tissue differs in its origin from that of the sexually produced plant, inasmuch 
as it arises as a simple strand from one of the numerous leaf-traces of the 
parent plant ; also, the earlier stages are hurried over or absent altogether, 
a fact which may be correlated with the relatively greater importance of 
the first leaf of the adventitious bud and with the early development of the 
commissural strand. Otherwise the stages in the elaboration of the stelar 
system are almost identical with those described above for the sporeling. 
The protostele at once becomes crescentic after the departure of the first 
leaf-trace ; the latter departs from the stem-stele as two strands which 
branch whilst still within the cortex of the stem. The trace of the first 
root fuses with the stem-stele opposite the point of departure of the first 
leaf-trace, whilst immediately above this point the commissural strand 
(Text-fig. 11, c.s.) arises by proliferation of the vascular tissues. A relatively 
