382 West. — A Contribution to the Study of the Marattiacea: 
times occurs, but the rhizome was unbranched in every specimen examined 
by the present writer. The numerous roots which arise from the ventral 
surface and flanks of the rhizome greatly outnumber the leaves, which form 
two ranks on the dorsal side of the rhizome. The leaves of Kaulfussia are 
not crowded, consequently distinct internodes can be distinguished (cf. the 
ill-nourished specimen of Danaea nodosa described above [p. 375] and cf. 
Text-fig. 14, A with 14, b). 
The development and arrangement of the vascular strands in the very 
young sporeling have been carefully studied by Campbell, who, in his 
monograph on the f Eusporangiatae ’, maintains that the vascular system 
of Kaulfussia is at first essentially similar to that of Danaea , consisting 
of a sympodium of the traces of the earlier leaves. He states, however, that 
Text-fig. 14. A. Danaea nodosa, Sm. Transverse section of the rhizome of an old (ill-nourished j 
specimen, x 5. B. Kaulfussia aesculifolia , Bl. Transverse section of the rhizome of an old 
specimen, x 4. c.s., commissural strand; r., root. 
the marked dorsiventrality of the adult plant has already impressed itself on 
the morphology and vascular anatomy of the young sporophyte, a distichous 
arrangement of the leaves being evident from the beginning, while the 
traces of the first and of all succeeding leaves are inserted upon the same 
side of the stelar system. Between the solid stele found in the very young 
stem and the dictyostelic cylinder of the older rhizome, Campbell (20, 
p. 184, Fig. 168, J, k) observed a transitional siphonostelic condition. 
The vascular anatomy of the adult plant of Kaulfussia has already 
been investigated by Kuhn (42), Farmer and Hill (29), and Gwynne- 
Vaughan (30). However, for comparison with that of the dorsiventral 
species of Danaea , the present writer examined the stelar system of the 
adult rhizome and petiole. 
