West. — A Contribution to the Study of the Marattiaceae. 389 
Text-fig. 18. Maraltia Cooperi , Mre. Model of the stelar system of the caudex of an adult 
plant, viewed from above. c.s.= commissural strand ; c.s.' = central cylinder ; comp.s. ■= compensating 
strand ; l.t., leaf-trace; r.t., root-trace (cf. Text-fig. 17). 
is probable that further complications 
arise in the stelar system of the huge 
tuberous stems of very old plants, 
but, unfortunately, no very old plant 
of Marattia was available for investi- 
gation. 
Vascidar A natomy of the Petiole . 
Gwynne-Vaughan ( 30 , pp» 264-5) 
found a large number of internal 
strands within the typical ring of 
bundles just above the basal pulvinus 
of a petiole of Marattia fraxinea ; 
traced upwards, these internal strands 
decreased in number by a succession 
cf fusions, which, however, take place 
so gradually that a number of the 
Text-fig. 19. Marattia Cooperi y Mre. 
Transverse section of the basal region of the 
stem of an old plant, showing numerous roots 
( r .) and leaf-traces (l.t.) embedded in the 
ground-tissue. The leaf-bases had been arti- 
ficially removed. The leaf-traces are numbered 
consecutively, l.t . 1 being the youngest shown. 
c.s. t commissural strand, x 1*5. 
