392 West. — A Contribution to the Study of the Marattiaceae. 
carefully elucidated ; the present writer’s investigations on this genus, 
although covering most of the ground, merely confirm previous results. 
Vascular Anatomy of the Petiole } The vascular anatomy of the 
petiole of Angiopteris appears to differ from that described above for 
Marattia only in the larger number of internal strands which arise from the 
strands of the abaxial arc ; as many as five separate concentric rings of 
bundles were counted by the author in a transverse section of the base of a 
petiole of an old plant. 
6. Macroglossum. 
This recently (discovered genus resembles Marattia and Angiopteris 
both in size and in the form of its upright, nearly globular, bulky stem, 
upon which the leaves are spirally arranged (cf. Copeland, 26 ; also cf. 
Campbell, 21 ). In the form of its simply pinnate leaves, however, it more 
closely resembles Danaea , 1 2 whilst its sporangia are very similar to those of 
A rchangiopteris. 
We are indebted to Campbell ( 21 ) for the only account of the anatomy 
of Macroglossum that has yet appeared. According to his account ( 1 . c., 
p. 66 1), no true cauline stele is developed in the young sporeling, the vascular 
system of the axis of which is at first composed only of leaf- and root- 
traces. A single root is formed for each of the early leaves. 
Sections of the basal region of the stem of a rather older specimen 
showed a type of stelar structure not far removed from that of the young 
sporophyte. In the centre there were five strands ; of these five strands, 
one was comparatively large and somewhat crescentic in transverse section, 
whilst the remaining four bundles, which were of smaller size, were arranged 
in pairs; the latter probably represent the double leaf-traces. 
It is to be regretted that the author ( 1 . c., pp. 662-3) only briefly 
refers to the anatomical features exhibited by the adult plant, which, as 
indicated above, unites within itself certain characters of four other genera 
of Marattiaceae. 
Apical Meristems. 
1. Stem . 
The structure and arrangement of the generative tissues at the apex of 
the massive stem and roots of the Marattiacean Ferns has been for many 
years the subject of much divergence of opinion. 
1 For an exhaustive account of the arrangement of the vascular strands in the petiole of 
Angiopteris , Marattia , and Kaulfussia , the reader is referred to Bertrand, C. E., et Cornaille, F. : 
Etudes sur quelques caracteristiques de la structure des Filicinees actuelles, in Trav. et Mem. de 
l’Universite de Lille, t. x, M^m. 29, 1902. 
2 Moreover the gametophytic characters of Macroglossum and Danaea agree in (1) the 
structure of the spermatozoids and in (2) the presence of a large suspensor (cf. Campbell, 21). 
