DEVELOPMENT OF CIEKIPEDIA. 
147 
They offer nothing at all peculiar, and consist of three joints, the first of which is the 
one from which the two branches start. Each of these consists accordingly of two 
joints. 
Between these natatory feet slender chitinous lamellae are seen running up on both 
sides of the abdomen (fig. 24, la), which separate the spaces from each other, in which 
after a short while the large cirri of the Lepas will develop, after the temporary feet of 
the Cypris have been deprived of the plastic substance which they contain. This 
shows out extremely well in one of my preparations, a Cypris which has just begun to 
give the first signs of metamorphosis ; also in fig. 25 this is still sufficiently visible. 
The two-jointed appendages of the tail are mostly turned upwards, and have setae on 
their last joints, similar to those on the natatory feet. 
The Organs of Sense. — Between the antennae, and a little behind them, two small 
protuberances might be observed, which Claus calls the “ Stirnhocker,” and which are 
very likely the remains of the lateral horns. No function seems, however, to be 
assigned to them now, as they have neither terminating glands nor any appendages 
which might be organs of touch. 
As sense-organs we have in the first place to claim the two olfactory hairs on the last 
joint of the antennae, and as organs of touch the setae and the peculiar elongate and 
flattened appendage which has been represented in fig. 27, u; also the setae on the 
sucker and the one on the second segment are very likely used as such. 
The nervous system is represented by two large ganglia (Plate 13. fig. 21, cer ), which 
are of an oval shape, and very likely united by a nervous bridge, which in our figure has 
been given with some doubt. These ganglia, in which nucleated cells may be observed, 
send large nerves to the base of the large compound eye, which consists of a black 
pigment body, eight to ten lenses and their cornea. The eye is at the bottom of a large 
capsule, very likely containing fluid, and attached to one of the valves by small muscles 
which enable it to move in different directions. Besides these large eyes we still find 
the Nauplius eye in the same place in which it was before, between the two antennae, 
a little above their points of attachment. 
Organs of Digestion and Glands. — The parts of the mouth which have been already 
described lead through a short oesophagus into the stomach, on both sides of which I 
saw, when dissecting the pupa, small glands, which seem to lead into the oesophagus 
as they do in the full-grown Lepas. 
The stomach is now rather collapsed, never distended by food like that of the 
Nauplius stage. It passes into an intestine, which is much longer than that of the 
latter, and ends most likely with an anus at the base of the last pair of natatory feet. 
I have, however, not been able to see this opening, nor could I find as yet a trace of the 
excretory caeca, which in the adult Lepas terminate on both sides of the rectum. 
Besides the oesophageal glands there are the cement-glands, which lead into the 
antennae and which have been traced from their first origin in the upper lip and 
described with the antennae. Very conspicuous as lying under the shell in the dorsal 
MDCCCLXXVI. 
X 
