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the Sporophylls of the Cycadaceae. 
while in the pinnae of the organ the bundles undergo a still 
greater reduction, and resemble there in their structure those 
of the ordinary foliage-leaf. 
Cycas circinalis, L. 
Female Sporophyll. In this plant the same structure of 
the strands prevails, but the latter are usually much elongated 
tangentially (Fig. 2). Some of these tend to become revoluted 
towards the dorsal side, a phenomenon similar to that oc- 
curring in the root of C. Seemanni , Al. Br. 1 
A single strand passes off to the sporangia, and in so 
doing divides into 3-5 strands which enter the latter ; some 
of those have a perfectly concentric structure. Fig. 3 repre- 
sents a bundle from the lamina in transverse section. 
The sporophylls are much more slender and the sporangia 
smaller than in the last species. 
Stangeria paradoxa, Th. Moore. 
Male Sporophyll. In the stalk are a number of bundles 
in a row (except at the very base where there are only two). 
In the well-developed centrifugal xylem the tracheides are 
clear and well defined, taking the stain brightly. Centripetal 
xylem is quite absent. In the sterile portion the centrifugal 
xylem is much reduced, its elements being inconspicuous 
and not so brightly stained as in those of the stalk ; in many 
bundles one, three, or four elements of centripetal xylem 
occur, in other bundles there are none seen at all. 
Female Sporophyll. Two bundles leave the cylinder of 
' the axis of the female cone, each from a separate strand, 
and may either enter the sporophyll without dividing further, 
furnishing to that organ two large bundles of equal develop- 
ment, or they may previously divide so as to form three 
or four bundles, of which the outer one on each side supplying 
the sporangium is considerably larger than the one or two 
1 Gregg, Ann. Bot., Vol. i, p. 4, Fig. i, 1887. 
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