446 
PROFESSOR BEALE’S NEW OBSERVATIONS UPON THE 
that their connexions were not so very distinctly seen, have been called “ connective- 
tissue corpuscles.” 
The drawings accompanying my paper explain the relation which I believe the essen- 
tial structures entering into the formation of the papilla bear to the indefinite con- 
nective tissue in which they lie imbedded. 
Epithelium. 
The so-called epithelium upon the summit of the papilla of the frog’s tongue (Plate 
XXI. fig. 1, a) differs from the epithelium attached to its sides ( b ), that covering the 
simple papillae (c), and that on the surface of the tongue generally, in many important 
characters. As is well known it is not ciliated. The cells differ from the ciliated cells 
in several points. They are smaller than these. The nucleus is very large in proportion 
to the entire cell. The cells are not easily separated from one another, as is the case with 
the ciliated epithelium. These cells form a compact mass, the upper surface of which 
is convex. This is adherent by its lower surface to the summit of the papilla, and it is 
not detached without employing force. The cells do not separate one by one, as occurs 
with the ordinary epithelium, but the whole collection is usually detached entire, and it 
is I believe torn away. 
Although some observers would assert that the two or three layers of cells repre- 
sented in my drawings do not exist, but that the appearance is produced by the cells of 
a single layer being pushed over one another by pressure, I am convinced that in this 
mass upon the summit of the papilla of the Hyla there is more than the single layer of 
cells represented by Hartmann, who is the latest observer on this point. 
Hartmann’s representation ( l . c.) of this very same structure from the summit of the 
papilla of the Hyla is very different from my drawings. Not only do we represent these 
same cells of very different shapes, but the nucleus in my specimens is three or four 
times as large in proportion to the cell as represented by him. 
The general outline of the free surface is convex (a, «, a, fig. 1, Plate XXI.), and 
the tissue which intervenes between the nuclei appears very transparent and projects 
a little, so as to give the convex summit a honeycombed appearance (Plate XXI. 
fig. 7). 
The under concave surface of this hemispheroidal mass which adheres to the summit 
of the papilla of the Hyla’s tongue, corresponds to the exact area over which the nerve- 
fibres of the papilla are distributed, as will presently be shown. The shape of these 
cell-like bodies, of which the mass is composed, and their connexion with fibres is shown 
in Plate XXI. fig. 3, and in the very highly magnified specimen represented in fig. 2. 
After the examination of a vast number of specimens I think these figures represent 
the actual arrangement, but this point is most difficult of investigation. In the inter- 
vals between what would be called, if they were capable of complete separation from one 
another, the individual cells, fibres are seen. These fibres do not I think arise simply 
from the pressure to which the masses have been subjected. I have represented the 
