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PROFESSOR BEALE’S NEW OBSERVATIONS UPON THE 
contain a small mass of germinal matter (nucleus) in the centre, which is of course 
coloured. In some of the smaller ones this mass of germinal matter is much larger in 
proportion to the entire “ cell.” These bodies resemble many of the fat-cells of the 
frog, and I think it probable they are of this nature. It is, however, possible that 
these masses may be altered lymph-corpuscles. The Hylae which I examined had been 
for some time in confinement, and contained very little adipose tissue. 
Conclusions. 
1. That fine nerve-fibres ramify in the connective tissue of which the simple papillae 
are composed, and that connected with these nerve-fibres are oval masses of germinal 
matter or nuclei, which are usually regarded as “ connective-tissue corpuscles.” 
2. That neither the epithelial cells of the frog’s tongue generally, nor those covering 
the simple papillae, are connected with nerve-fibres. 
3. That the mass consisting of epithelium-like cells upon the summit of the fungiform 
papilla, is connected with the nerve-fibres, but it is not an epithelial structure. 
4. That the dark-bordered sensitive fibres constituting the bundle of nerves in the 
axis of the papilla divide near its summit into numerous very fine branches, with which 
nuclei are connected. Thus is formed a plexus or network of exceedingly fine fibres 
upon the summit of each papilla ; from this network numerous fine fibres may be traced 
into the special nervous organ, composed of epithelium-like cells upon the summit, with 
every one of which nerve-fibres appear to be connected. 
5. That the bundle of nerve-fibres distributed to a papilla always divides into two 
bundles which pursue opposite directions. The division of the bundle may take place 
just at the base of the papilla, or at some distance from it, but it always occurs. 
6. That fine pale nerve-fibres pass from the same trunk of dark-bordered fibres as that 
which gives off the bundle of nerves to the papilla. The fine fibres ramify — 
a. Amongst the muscular fibres of the tongue. 
b. Upon the vessels. 
c. In the connective tissue of the tongue generally, and also in the simple papillae. 
7. That the fine nucleated nerve-fibres ramify freely amongst the delicate branching 
muscular fibres of the papillae, and form plexuses or networks which exhibit no nerve- 
ends or terminations, nor in any case does a nerve-fibre penetrate through the sarco- 
lemma or investing tissue of the fibre, or connect itself with the nuclei of the muscle. 
As many of the muscular fibres are so very fine and narrow, the distribution of the nerves, 
and their exact relation to the contractile tissue, can be demonstrated very distinctly in 
the case of the muscles of the papillae of the frog’s tongue. 
