456 
DR. HERBERT WATNEY ON THE MINUTE 
injected into the lymphatics of the villi left the vessel and passed into the stroma of the 
villus (see Taf. iii. fig. ii.). He showed that the walls of the chyle-vessels are lined by 
an endothelium. Recklinghausen thought there was a Saftcanalsystem in the villi, 
although he allowed that there is a gradual transition from follicular tissue (adenoid 
tissue, His) to the Saftcanalsystem. 
His (51) and Auerbach (52) confirmed Recklinghausen’s observation of the existence 
of an endothelial covering to the chyle-vessels. His maintained with Teichman that the 
injection does not leave the chyle-vessels. Notwithstanding these researches, three 
observers, Fles (53), Basch (54), and Zawarykin (56), declared that in its upper part 
the central vessel has no definite membrane. 
Basch (54), 1865, found no difference between the cells of the villus and those lining 
the chyle-vessel. He injected from the lymphatics, and not only filled the central canal 
of the villus, but his injection penetrated the stroma of the villus in lines, forming a 
regular network. 
Letzerich (55), 1866 and 1867, described certain structures among the epithelial 
cells, which in the normal state are the sole absorbents of fat; these structures 
(“ vacuoles ”) are connected with a system of canals, and the canals with the central 
vessel. In pathological states the epithelial cells contain fat. When not in action the 
vacuoles are closed, and are seen as star-shaped bodies between the epithelium. Eimer 
and other observers pointed out that the vacuoles are “ goblet-cells.” 
Kolliker (44), 1867, confirmed Eberth’s researches as to the basement membrane. 
He held, however, that the border seam is nothing but a thickened outer layer of the 
connective tissue, corresponding to the border layer of the “ follicular glands.” Kol- 
liker considered it probable that the fat passes through the pores of the epithelial 
cells ; and stated that the paths by which the fat travels in the tissue of the villus are 
unknown to anatomists. 
Erdmann (34), 1867, found that the epithelium is separated from the stroma of the 
villus by an unbroken membrane. This membrane sends processes, on the one hand, 
around the epithelial cells (as a cement substance), and, on the other hand, in the stroma 
of the villus (as connective-tissue trabeculae)*. These trabeculae enclose the connective- 
tissue corpuscles. The corpuscles of the villus are not star-like, but round ; the fat 
travels by the cement substance between the epithelial cells. 
Eimer (41), 1869, tried to demonstrate Heidenhain’s theory. He gave drawings of 
the connexions of the epithelial cells with the tissue of the villus from sections of a 
hardened intestine of a frogf. Eimer further gave drawings of fat-particles in the pro- 
cesses of the branched connective-tissue corpuscles of the submucosa of the frog’s intes- 
* Eies (1865) considered that the intercellular substance between the epithelial cells is in direct communi- 
cation with the connective-tissue of the villus. 
t Almost all the attempts to establish the connexions of the epithelial cells and the connective-tissue cor- 
puscles have been made from preparations of the frog’s intestine ; and this is the more remarkable from the 
fact that the epithelial cells in that animal are very long and narrow, and that it is somewhat difficult to show 
their connexions. 
