284 Ewart — On the Leaf -glands of 
was taken from a section of a fresh leaf which had not been in 
alcohol, but had been simply cut and mounted in water. The 
whole gland was very similar to that shown in Fig. 5. Some 
transverse walls had appeared at intervals in the secretory 
cells, but the majority of them were undivided, and very long 
and narrow. The nuclei, still large but not so conspicuous, 
occupied various positions in the cells, but for the most part 
were situated in the basal or middle portions. 
The protoplasm had become much vacuolated, and this 
gave the cells a lighter, somewhat swollen, appearance. The 
two supporting cells below each group of secreting cells con- 
tained little protoplasm, and the lateral walls were thickened 
and cuticularized to a considerable extent, especially in the 
upper ( has 1 ). Fig. 17 represents a section taken across a 
group of epithelium-cells. 
While cutting some of the smaller leaves, I noticed a large 
number of small apparently sessile glands scattered over the 
surface both of petiole and blade (gl. Fig. 6). Examination 
under a higher power showed that each was in reality shortly 
stipitate, being supported on a single cell. The epidermal 
cell below this stalk was slightly depressed, so that the cluster 
of four or five cells constituting the gland proper appeared 
to rest on the epidermis (Fig. 19). 
In rather older leaves the glands were separated by wider 
intervals, and the depression of the epidermis round each 
gland was more marked (Fig. 20). 
The secretory cells had multiplied by divisions vertical 
to the surface, but the stalk-cell had undergone very little 
alteration. The wall dividing it from the epidermal cell 
immediately below had become thickened and cuticularized, 
and now the resemblance to an early stage of a single bundle 
of secretory epithelial cells from the large glands already 
described, with its two basal cells, became obvious. Even 
in the youngest leaves, before there was any trace of the 
invagination to form the future large gland, these small 
surface-glands were partially formed, and consisted usually 
of epidermal cell (bas l ) } stalk-cell (bas 2 ), and group of two 
