294 Phillips .- — On the Development of 
the terminal filament (Figs. 2, 3, 5, 7). The curvature of the 
carpogonial branch is such that, as has been described by 
Schmitz, the carpogonium itself lies medianly above and close 
to the pericentral cell, from which the carpogonial branch 
arose. This pericentral cell was first regarded by Schmitz as 
the auxiliary cell, but he now regards a later derivative of it, 
which had previously escaped his notice, as the true auxiliary 
cell ( 7 ). There is, however, some reason for adhering to his 
first statement, and I will return to this point again, de- 
scribing the pericentral cell as the auxiliary for the present. 
The carpogonial branch does not occupy a definite position 
in the young cystocarp right or left of the two-celled lateral 
branch, from which it may be inferred, I think, that this lateral 
branch is the morphological equivalent of the carpogonial 
branch. 
The derivatives of the pericentral (auxiliary) cell at this 
stage consist therefore of the following : — 
(a) A four-celled carpogonial branch, ending in the carpo- 
gonium with its trichogyne ; 
(b) A two-celled sterile lateral branch facing it ; 
(c) A one-celled sterile inferior branch. 
In the majority of the numerous procarps which are crowded 
on the tufted branches of this plant, no fertilization takes 
place. The procarps do not, however, thereupon atrophy and 
disappear, but, as before stated, are gradually metamorphosed 
into vegetative shoots. The plant thus offers a convenient 
opportunity of sharply distinguishing between the condition 
of the procarp before fertilization and the developments 
which are consequent upon fertilization. With regard to the 
proliferation itself, it is not clear how it is effected, what cell 
for example of the former procarp becomes the leading-cell of 
the branch. Two points are clear, however. First, that no 
cell of the axial row becomes the apical cell, so that the proli- 
ferated branch is a sympodium ; secondly, that all the cells of 
the former procarp become active, living, cells of the branch, 
excepting only the four cells of the carpogonial branch. The 
carpogonium itself on the failure of fertilization becomes ill- 
