Notes. 
667 
(1) As the spore-mother-cell approaches the period of division, the 
nucleus becomes oval, and the linin aggregates in the vicinity of the 
nucleolus, which is commonly situated at or near one end of the 
nucleus. Some portions of the linin however still protrude as threads 
or network from the aggregated portion. 
(2) The nucleolus suddenly becomes vacuolated and often frag- 
ments, the linin is seen to consist of a tangle of double filaments, 
indicating that the longitudinal fission has already occurred. Frequent 
anastomoses are obvious. 
(3) The nucleus becomes asymmetrical owing to the linin and 
nucleoli becoming massed about the middle of one side. 
(4) The achromatic spindle next appears with extraordinary sudden- 
ness. It is bipolar, but of a peculiar form. The polar angles of the 
spindle are very wide, whilst the depth of the flat spore-mother-cell 
is comparatively slight. Thus the transverse section of the spindle 
at the equator becomes elliptical instead of circular, and the chromo- 
somes are seen to be collected on one side, instead of being uniformly 
distributed around the equator (cf. par. 3). When observed from the 
flat side of the cell, the spindle is seen to present the form of 
a triangle, at two of the angles of which fine centrospheres are visible, 
whilst the chromosomes are grouped at the remaining angle. It must 
be stated that this appearance is seen both in sections, and also in un- 
injured cells teased out of the sporogonium and mounted in glycerine 
tinged with eosin. The general effect produced strongly recalls the 
development of the spindle during the division of the spermatocytes 
of Salamanders, as described by Hermann 1 , and indeed his figures 
8 and 9 would almost serve to illustrate these spindles in Fegatella. 
The origin of the structure is, however, different in the two cases, and 
the form in Fegatella is probably connected partly with the contour of 
the cell, and partly with the existence of the lateral aggregation 
already (3) referred to. The strand connecting the two centrospheres 
consists of unbroken threads, and it is weaker than those which run 
from these bodies to the chromosomes. 
(5) Very extensive radiations extend from the centrospheres into 
the surrounding protoplasm, and they may often be traced quite to the 
periphery of the cell. Furthermore, weak radiations also extend from 
the equatorial patch of chromosomes towards the nearest wall. The 
latter filaments suggest the existence of an early tripolar spindle ; and 
1 Hermann, Archiv fiir mikr. Anat., Bd. xxxvii. 
