Observations on the Development of 
Marattia Douglasii, Baker. 
BY 
DOUGLAS HOUGHTON CAMPBELL. 
Professor of Botany, Leland Stanford Junior University , California, US. A. 
With Plates I and II. 
HILE collecting in the Hawaiian Islands during the 
V V summer of 1892, I was fortunate enough to find near 
Hanalei, upon the island of Kauai, a large number of very 
young plants, of Marattia Douglasii, and with them a suf- 
ficient number of prothallia with embryos to make it possible 
to study the principal points in their development. Until 
Farmer’s 1 recent paper on Angiopteris , no account of the 
embryogeny of the Marattiaceae has appeared, except a brief 
mention made by Luerssen 2 of the arrangement of the 
primary organs. His material was however too scanty to 
throw any light upon the early divisions of the embryo. The 
growth of the prothallium is very slow and the development 
of the embryo correspondingly late, so that in artificial 
cultures more than a year must ordinarily elapse before 
embryos are developed. This probably accounts for the fact 
1 Farmer, On the embryogeny of Angiopteris evecta, Annals of Botany, Vol. vi, 
No. XXIII, Oct. 1892. 
2 Luerssen, Handbuch der Systematischen Botanik, Vol. i, p. 582. 
[Annals of Botany, Vol. VIII. No. XXIX. March, 1894.] 
