NOTES. 
THE SIEVE-TUBES OF CALYCANTHUS OCCIDENT ALIS 
(Hook and Arn.) — In Calycanthus , De Bary describes the four cortical 
bundles with reversed orientation, and with regard to their phloem 
he remarks, ‘ It only consists of soft bast, and, in the main at least, 
of parenchymatous elements; sieve-tubes remain still to be sought 
for 1 / Herail, who subsequently studied the origin and development 
of these bundles, states 2 that instead of being cortical they are derived 
from the pericycle. The hypocotyledonary stem is described as 
possessing four masses of phloem, but no cortical bundles. The 
latter become separated from the phloem of the bundle-ring about 
the middle of the first internode. Herail states that sieve-tubes are 
present in this part of the stem, but he makes no mention of them 
in connexion with the cortical bundles. 
Young stems of C. occidentalis were examined and the greater 
part of the phloem in the cortical bundles was seen to consist of 
sieve-tubes. On comparing the bast of these bundles with that of 
the bundle-ring it was found, as a matter of fact, that, for equal areas, 
the sieve-tubes in the former were far more numerous than in the 
latter. 
The following details may be added. A transverse section shows 
the cortical (pericyclic, according to Herail) bundle embedded in 
the cortex, and separated from the normal phloem by a very variable 
number of parenchymatous cells with large interspaces, see woodcut p x . 
Its shape is that of a sector of a circle with the angle towards the 
periphery, and the arc near the bundle- ring, while the radii diverge 
at an angle of about 120°. A semilunar band of sclerenchymatous 
1 De Bary, Comp. Anat. Eng. Ed. p. 584. 
2 Herail, * Recherches sur l’anat. Comp, de la Tige des Dicotyledones/ Ann. 
Sci. Nat. Bot. 7 Ser. Tome 2, p. 236. 
