58 Eriksson . — On the vegetative life of some Uredineae . 
principal outbreak in the new year can take place in the beginning of June 
or even earlier, in the course of May. Thus in the past season 1903 I have 
seen on November 2 pustules of llredo glumarum abundantly in several 
wheat-plots on the experimental- field, and new pustules were already 
breaking out in great abundance in the first days of May. In such a season 
the period lacking mycelium in the wheat-plant’s life must be short, two 
or three weeks in October after the sprouting, and in the new year, two or 
three months, March-May, in the early spring. 
III. The Intracellular Mycoplasm-Life of the Fungus. 
The results of the researches carried out have, however, by no means 
been only negative. In all embeddings of varieties especially susceptible 
to the disease we find some cells more or less filled tip with a particidar 
dense plasmatic substance , at first commonly fine-grained, later more reti- 
form and vacuolar. In the younger stages — the resting-stage of the my co- 
plasm — the cell-nuclei seem to be of normal size, but later they swell 
to twice their normal size and lose to a certain degree their normal fibrillar 
structure. Gradually they also lose their normal form and appear as if 
eroded on the surface. At last they usually disappear as organized bodies, 
vacuoles frequently appearing as well as some small bodies, which take the 
saffranin stain. Now the mycoplasm enters into its ripening -stage. 
From this time the Fungus must be considered as a true parasite, 
beginning to dissolve the cell-nucleus. Simultaneously one sees in the 
mycoplasmic reticulum small spherical bodies, which take the saffranin 
stain. These bodies are surrounded by hyaline areas and are believed 
to be the nucleoli of the nuclei of the mycoplasm , the hyaline area sur- 
rounding each nucleolus being the body of the nucleus. The spherical 
bodies are very variable in size. 
IV. The Intercellular Mycelium-Life of the Fungus. 
Now the plasmodium is ready to force itself out of the host-cell, 
in order to develop an intercellular mycelium. On its forcing itself out no 
dissolution of the cell-wall, either total or partial, takes place. It seems to 
be the case. that the plasmodium effuses through the subtile pores that 
must be supposed to exist in the cell-wall, that is to say in the same 
way as the plasmodesms between the cells. 
At this stage of development we often find places in the preparations, 
where wholly corresponding portions of plasma are to be found inside 
and outside the cell-wall. The connexion between the two plasma-portions 
is only broken off in consequence of the plasmolysis of the cell-contents. 
