1 1 8 Sargant and Robertson . — The Anatomy of the 
washed out by the water in which the tissues of the young seedling are 
soaked, and perhaps the internal tissues which lie round the gland have 
swollen, closing its narrow cleft and expelling the secretion at its mouth. 
Such swelling of internal tissues may also account for the comparative 
absence of wrinkles in the dorsal surface of the scutellum belonging to 
a growing embryo. In such embryos many glands open out on a smooth 
layer of epithelium, whereas in the dry seed we cut they nearly always 
started from the base of a depression. 
The glands vary greatly in size. Some of the smallest are funnel- 
shaped pits, others shallow slits, and it is very difficult to draw the line 
between structures which deserve the name of glands and mere depressions 
or wrinkles. The largest we measured were clefts of considerable size. 
The opening of one was *66 mm. long, and its maximum depth -i mm. 
Another reached a depth of 8 mm. 
The number of glands found in a single scutellum is also very variable. 
Three sets of our serial sections include the whole of the scutellum ; and 
we have counted the number of glands in each series. The numbers thus 
obtained have no great absolute value. Two difficulties stand in the way 
of exact determination. The line between well-marked depressions and 
small glands is not easily drawn, and glands which run nearly parallel 
to the plane of section are readily passed over. The numbers recorded 
are therefore in all probability too small. 
In seedling A 9 (Fig. 9, PI. V), six days old, the scutellum was cut 
transversely. Thirty-eight glands were counted. 
In seedling a, two days old, the scutellum was cut transversely. Only 
seven glands were found, all small. This is probably an exceptional case. 
In seedling b> two days old, in which the scutellum was cut longi- 
tudinally, twenty-nine glands were counted. 
Sections were also cut transversely through an imperfect scutellum, 
dissected out of an ungerminated seed. The extreme tip and extreme 
base were absent, but thirty-eight glands were counted in the remainder. 
The majority of glands counted in those seedlings which were cut 
transversely run parallel with the longer axis of the scutellum, while in 
seedling b, which was cut longitudinally, the majority are more or less 
perpendicular to the longer axis. In other words, more glands were always 
counted in the direction perpendicular to the plane of section, and this 
result is clearly due to the difficulty of recognizing a gland which lies 
parallel to the section. But we believe that longitudinal glands are really 
rather more common than transverse ones. 
The glands are scattered over the whole surface of the epithelium 
except where it covers the top of the scutellum. In seedlings A 9 , a, and b } 
no glands are found above an imaginary line drawn across the scutellum 
at a distance from the apex of one-third its whole length. The apex of 
