Mottier . — The Embryology of some Anomalous Dicotyledons. 455 
denary primordium appears in the form of a narrow, elongated depression, 
which is just perceptible at the truncated end, extending from one side 
to the centre. The shaded part of b, Fig. 29, represents this depression. 
The primordium here is more nearly kidney-shaped than crescentic. 
Fig. 11 represents a more detailed drawing of a , Fig. 29. The larger 
cells in the centre indicate the apex of the stem. There is at this stage 
no indication of the plerome strands; these appear later, as shown for 
the species already described. 
As growth continues, the primordium becomes typically crescentic, 
and the embryo is more cylindrical (Fig. 30, a-e , representing alternate 
sections of a part of the embryo). Whether the embryo represented 
in this figure was not cylindrical from the beginning cannot be stated. 
However, this matter is unimportant. During succeeding development, 
the crescentic primordium soon bifurcates, but the difference in depth 
of the two sinuses is increased rather than diminished, so that in the 
mature seed the anomaly is more pronounced (Fig. 31, a-g). This figure 
does not include all of the embryo beyond the base of the cotyledons, 
but only alternate sections from that point to the bottom of the second 
sinus. This series of sections, taken from an embryo in the mature 
seed, shows that a cotyledonary tube is present, a phenomenon that has 
been reported for two other species of Delphinium , D. nudicaule , and 
D. hybridum (Sargant, ’ 03 , p. 73). Fig. 32 is intended to show the 
form of the embryo as it is in the mature seed. As it was found 
impracticable to dissect out the embryo on account of its small size and 
the firmness of the endosperm, this figure was constructed, as in the 
other species, from a complete series of cross-sections, and the proportions 
are reasonably accurate. It may be noted that the embryo has a relatively 
longer hypocotyl than in Actea , and the cotyledons diverge towards 
the apex. These are crescentic in cross-section. The two sinuses are 
of unequal depth, and the cotyledonary tube is formed by the intercalary 
growth of the common base of the cotyledons. In Fig. 32, the relative 
depth of this tube is indicated by the dotted line, which shows also the 
outline of the space enclosed by the cotyledons. In Delphinium , as 
well as in the other species under consideration, there is no plumule 
in the ordinary conception of that term. Here the apex of the stem 
does not even project as a recognizable protuberance, at least, so far as my 
observations have extended. 
Aquilegia canadensis. 
The youngest stage of the embryo of Aquilegia observed was an 
elongated cylindrical structure, somewhat bent near the middle, and 
possessing a suspensor of three or four cells (Fig. 12). At this stage, 
it was not possible to determine the exact limits of the suspensor. With 
