A Study of the Vascular System of Matonia 
pectinata. 
BY 
A. G. TANSLEY, M.A., 
Assistant-Professor of Botany, University College , London , 
AND 
MISS R. B. J. LULHAM, B.Sc. 
With Plates XXXI, XXXII, and XXXIII, and five Figures in the Text. 
Contents. 
PAGE 
Introductory 475 
Young Plants 476 
The Morphology of the Leaf 477 
Details of Progressive Complication in the Vascular System : — 
(1) Young Plants 482 
(2) Intermediate Plants ............. 490 
(3) Adult Plants 492 
Summary of Progressive Complication in the Vascular System 496 
Protoxylems 503 
Nature and Phylogenetic Relations of the Vascular System in Matonia : — 
Morphological Position and Origin of the Matonia-type 508 
Functional Relations of the Vascular System of Matonia in connexion with 
its Evolution 510 
Roots . 513 
Morphological Status of ‘ Pith ’ . 514 
Introductory. 
Ever since the meeting of the British Association at Bristol in 1898, 
at which Mr. Seward described 1 the complicated and striking vascular 
anatomy of the comparatively rare Malayan fern Matonia pectinata , the 
problem of relating this apparently unique vascular skeleton to those of 
other Ferns has been a matter of considerable interest to students of 
Filicinean vascular morphology. 
1 Subsequently published in full, Phil. Trans. B., vol. cxci, p. 171, 1899. 
[Annals of Botany, Vol. XIX. No. LXXVI. October, 1905.] 
