Notes. 
1 7 1 
superior (ventral or posterior) and three to the inferior (dorsal or 
anterior) lobe. 
The segments are of two kinds — sterile and fertile. Both alike 
consist of a long, straight, slender pedicel, running out horizontally, 
and terminating at the distal end in a thick laminar expansion. 
The sterile segments are the longer, and their laminae bear, in 
each case, an upturned foliaceous scale as well as a shorter and 
stouter downward prolongation 1 . 
Each of the fertile segments ends in a fleshy laminar enlargement 
not unlike the peltate scale of an Equisetum or a Calamostachys . 
These fertile laminae, which are protected on the exterior by the 
overlapping ends of the sterile segments, bear the sporangia. Four, 
perhaps in -some cases five, sporangia are attached, by their ends 
remote from the axis, to the inner surface of the peltate fertile lamina. 
Each sporangium is connected with the lamina by a somewhat narrow 
neck of tissue into which a vascular bundle enters. The sporangia 
are of great length, and extend back along the pedicels until they 
nearly or quite reach the axis. 
The sterile and fertile segments alternate regularly, one above the 
other, in the same vertical series. So much is evident, but the 
question, which segments are fertile and which sterile, has presented 
great difficulties, owing to the fact that the same segment can scarcely 
ever be traced continuously throughout the whole of its long course, 
and that the pedicels of sterile and fertile segments present no 
constant distinctive characters. For reasons, however, which will be 
fully given in a subsequent paper, I think it highly probable that in 
each sporophyll the segments of the lower lobe are sterile, and those 
of the upper lobe fertile, constituting the sporangiophores. 
The sporangia and pedicels are all packed closely together so as to 
form a continuous mass. The external surface of the cone was com- 
pletely protected by its double investiture of fertile and sterile laminae. 
The spores are well preserved in various parts of the cone, and, so 
far as this specimen shows, are all of one kind, their average diameter, 
being 0-065 mm - At the base of the cone, where macrospores, if 
they existed, might naturally be looked for, the spores are of the same 
size as elsewhere. So far, then, there is no evidence of heterospory. 
1 I now find that both the upturned scale and the downward prolongation are 
in reality double. D. H. S., February 11, 1897. 
