The Formation of the Sexual Nuclei in 
Lilium Martagon : 
II. Spermatogenesis. 
BY 
ETHEL SARGANT. 
With Plates X and XI. 
I N the first part of this paper I described the three nuclear 
divisions which take place within the embryo-sac of 
Lilium Martagon and form the nucleus of the ovum there. 
The reasons which led me to undertake a fresh examination 
of material rendered classical by previous research will be 
found at the beginning of that part. My object was to 
examine each of those three nuclear generations with the 
single aim of determining whether there was ground for 
the belief that any one of the nuclei in that series was formed 
of chromosomes derived from those of the previous generation 
by transverse fission. Dr. Haecker’s hypothesis, which I have 
discussed elsewhere (I, pp. 445-9 1 ), demands that one of the 
three nuclei in question should be so formed 2 . It further 
demands that in the parallel series of four nuclear generations 
which terminates in the formation of the male pronucleus 
1 This and all similar references are to Part I of this paper, Annals of Botany, 
x, 1896. 
2 Dr. V. Haecker, The Reduction of the Chromosomes in the Sexual Cells as 
described by Botanists, Annals of Botany, ix, 1895. 
[Annals of Botany, Vol. XI. No. XLII. June, 180'?.! 
