350 Phillips . — On the Development of 
it is deflected towards the axis by the more rapid growth of 
the external peripheral cells of that joint. The proximal cell 
of the row of peripheral cells thus arising on the fifth joint, 
now begins to give off cells in all planes, which are bound 
together in a common mucilaginous mass and constitute 
a small cup. The deflected axial row shares, on the internal 
(ventral) side, in the formation of this cup. The second cell 
of the external peripheral row now gives rise above to 
a two-celled carpogonial branch, which occupies the hollow of 
the cup and extends beyond in an unusually short trichogyne 
(PI. XVII, Fig. i). A considerable proportion of the procarps 
which reach this stage do not develop further, presumably 
through failure of fertilization. In those which do, two things 
soon become marked. First, the rapid growth of the fila- 
ments constituting the cup, until this body becomes urn- 
shaped with an apical pore. The lower cells cut off, externally, 
cells which form a cortex to the pericarp, continuous with the 
cortex of the stalk. Secondly, the interior of the urn becomes 
filled with a compact hemispherical mass of tissue, the cells 
of which radiate from the base upwards and are of a bright- 
red colour (PI. XVII, Fig. 2). At first, I considered this 
tissue to be the gonimoblast-filaments originating in an 
auxiliary cell. I discovered however that they gradually 
disappeared again in successively older cystocarps, and that 
the real gonimoblast-filaments arose much later, and consisted 
of larger and stronger filaments. These true gonimoblast- 
filaments surround the first crop of sterile filaments (PI. XVII, 
Fig. 3), which become gradually more and more attenuated, 
until in maturer cystocarps they could not be traced. Re- 
verting to the origin of the sterile filaments, I found, to the 
best of my belief, that they arise from the first cell of the 
carpogonial branch, the cell, that is, which lies directly 
beneath the carpogoniurri. By their luxuriant growth, the 
carpogonium itself is thrown over upon the base of the cavity 
of the cystocarp, where it may be seen at this stage, dis- 
tinguishable only by its mucilaginous wall running out into 
the remains of the trichogyne. The origin of the gonimoblast- 
