Notes. 
59i 
it first Lepidostrobus insignis , and afterwards Lepidodendron Spenceri 
while the other ( Spencerites majusculus ) is new. 
In one of his latest publications, Williamson pointed out that it 
might ultimately be necessary to make his Lepidodendron Spenceri the 
type of a new genus 1 2 . The separation thus suggested is now carried 
out, on the basis of a renewed investigation of the structure of this 
fossil. 
Spencerites insignis is a pedunculate strobilus ; the vegetative organs 
are not as yet identified. The specimens are calcified, and their 
structure admirably preserved. 
The anatomy of the axis is of a simple Lycopodiaceous type, but 
differs in details (such as the course of the leaf-trace bundles) from 
that of the axis of Lepidostrobus. The peduncle bears sterile bracts, 
similar to the sporophylls of the cone itself ; the latter are arranged 
spirally, or in some cases in alternating verticils. 
The individual sporophylls are of peltate form, consisting of a short 
cylindrical pedicel, expanding into a relatively large lamina. The 
sporangia are approximately spherical bodies; unlike those of Lepido - 
strobus , they are quite free from the pedicel, and are attached by 
a narrow base to the upper surface of the lamina, where it begins 
to expand. 
The details of the sporangial wall are quite different from those of 
Lepidostrobus , and the spores are also characteristic. In size they 
are intermediate between the microspores and macrospores of 
Lepidostrobus. They are of tetrahedral form, becoming spheroidal 
when mature, and each spore has a hollow, annular wing running 
round its equator. The wing is no doubt formed by a dilation of 
the cuticle 3 , and not, as Williamson supposed, from the abortive 
sister-cells. 
Spencerites majusculus , the new species, is much larger than the 
former, the axis of the cone being twice as thick. The anatomy is 
similar, but the sporophylls, and consequently the leaf-traces, are 
more numerous. The sporophylls, which are arranged in alter- 
nating verticils, are relatively short, and of peculiar form ; the lamina 
is very thick, and of great tangential width. The sporangia are like 
1 Williamson, Organization of the Fossil Plants of the Coal-measures, Parts 
IX, X, XVI, and XIX, Phil. Trans., 1878, 1880, 1889, an( l 1893* 
2 General Index, Part II, 1893, p. 24. 
3 Cf. Solms-Laubach, Fossil Botany, p. 239. 
