EEPOET ON THE EXPLORATION OE BEIXHAM CAVE. 
531 
point out any characters, either in size or form, sufficient to distinguish the Spelaean 
from the Spotted Hyaena. 
In the subjoined Table I have given the mean, maximum, and minimum dimensions 
of the principal teeth in different species of Hyaena, from which it will be seen how very 
closely, in almost every tooth, the fossil Hyaena agrees with II. crocuta ( fera ), or, rather, 
as I should explain, with the larger variety of that species. Taking the individual teeth, 
it would seem that the upper canine is somewhat larger in the existing Hyaena, whilst 
the third premolar is a trifle less, its maximum size only coming up to the mean in 
Hycena spelcea. The fourth upper premolar is considerably less, even its maximum size 
falling considerably short of the mean in the fossil form. The lower canine, again, like 
the upper, is somewhat larger in the existing species, though the difference is very 
slight. The first lower premolars are of nearly equal size in both, as are also the 
second and third ; but the lower carnassial tooth appears to be sometimes a good deal 
larger in the fossil Hyaena, although its mean dimensions are but very little more. 
Table showing the Mean, Maximum, and Minimum Dimensions of certain Teeth 
in different Species of Hycena. (0"-01.) 
Canine. 
pm 3. 
pm 4. 
Canine. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
H. crocuta (fera)* ... 
76x50 
80x50 
55x40 
94x70 
100x70 
71X60 
146x84 
150x85 
130x75 
66x49 
75x50 
55x40 
H. brunnca 
67X50 
70x52 
65x50 
93x63 
100x67 
90x60 
142x85 
150x90 
140x82 
72x50 
75x50 
70x50 
H. striata 
60x40 
70x40 
55x40 
80x52 
90x60 
75x51 
117x70 
120x77 
110x66 
54x41 
60x40 
50x40 
H. spelcea 
74x46 
100x73 
160x87 
71 x52 
H. (Brixham) 
70x51 
70x55 
70x50 
100x70 
160x90 
62x50 
65x50 
60x50 
pm 1. 
pm 2. 
pm 3. 
m. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
Mean. 
Max. 
Min. 
H. crocuta (fera) * ... 
63x44 
70x48 
50x35 
85x60 
90X62 
70x50 
91x52 
95x50 
70X50 
120x50 
120x55 
100x40 
II. brunnea 
62x45 
70x50 
60x45 
84x55 
90x60 
80x52 
94x53 
95x55 
95x53 
94x58 
97x50 
95x45 
H. striata 
52x32 
60x35 
47x30 
72x44 
75x48 
70x40 
78x43 
80x45 
72x40 
81x40 
85x40 
77 X40 
H. spelcea 
66x48 
89x65 
95x59 
128x54 
H. (Brixham) 
61x46 
65x48 
60x45 
90x61 
90x65 
90X60 
92x5S 
95x60 
90x55 
125x53 
132x52 
120x50 
On the whole, when we consider the fact that the dental differences between H. 
crocuta and H. spelcea are to the full as great, if not greater, than they are between the 
Lion and Tiger, I am hardly at present inclined to agree with Messrs. Boyd Dawkins 
* The minimum measures under II. crocuta are taken from caged specimens. There is no reason for sup- 
posing that the teeth are ever so small in the •wild state. 
