Biology of Fegatella conic a . 109 
abundant starch-grains. In the upper portion of the sporogonium there is 
evident at a relatively early stage a well-marked outer layer which gives 
rise to the wall of the capsule, the inner cells forming the archesporium , 
from which arise the spores and elaters. 
Spores. 
The archesporial cells are at first all alike, but later they become 
differentiated into two kinds, which show no definite relation to each other 
Fig. 32. I. Venter of fertilized archegonium, with eight-celled embryo (octant-stage), four 
cells being seen in the section. II. Transverse section of a similar embryo. Ill, IV. Older 
embryos in longitudinal section. V. Part of longitudinal section of developing capsule, showing 
differentiation of archesporial tissue into long narrow elater-forming cells (el.) and nearly isodiametric 
sporogenous cells (sp.). VI. Part of longitudinal section of receptacle, showing a young sporogonium 
in outline. VII. Similar section, showing two mature sporogonia in outline, a. c. air-chambers ; 
cal. calyptra ; caps, capsule ; f. foot ; n. withered neck of archegonium ; p. pore ; rec. si. stalk of 
receptacle ; rhiz. rhizoids ; s. seta of sporogonium. I-V, x 360 ; VI, x 76 ; VII, x 20. 
beyond a tendency to become arranged in longitudinal rows (Fig. 33, V). 
Some of the cells grow in length and remain narrow, whilst the others 
grow almost equally in all directions, and are further distinguished from 
the long cells by containing more densely granular protoplasm. The 
larger cells eventually become rounded off and constitute the spore mother- 
cells. The nucleus divides into two, and then each daughter-nucleus divides 
