172 Benson. — Telangium Scotti , a new Species of 
for our purpose to refer to the drawings of Scolecopteris polymorpha in 
Engler and Prantl, Teil I, Abt. 4, p. 440. It will be seen that the sorus 
as a whole somewhat resembles T. Scotti , but the four or five sporangia, 
which here constitute the sorus, are inserted around a pedicel along which 
runs a vascular strand. If this were to become continuous with a strand 
of trachei'des developed in the sporogenous tissue, we should obtain the 
vascular supply which characterizes Lagenostoma. 
The beautiful plates in Brongniart’s ‘ Recherches sur les graines 
fossiles silicifiees 5 afford many opportunities of applying and testing 
the new theory, and amongst others I would suggest a reference to the 
following : — 
Plate IX, Fig. 4, showing a vascular bundle entering the nucellus in 
Rhabdocarpus subtunicatus. 
Plate XIII, Figs. 6 , 7, 17, 19, showing the contrasted tissue-systems of 
the integument of Sarcotaxus avellana and its septicidal dehiscence. 
Plate IV, Figs. 1 and 3, showing the sporangial appearance of the 
inner integument continued to the base of the nucellus in Cyclocarpus 
nummularis. (These figures should be compared with Telangium Scotti , 
Fig. 8.) 
Plate C, Fig. 9, in which the seed Codonospermum is shown to present 
a striking external resemblance to such a synangium as Asterotheca . 
The similarity of the inner integument of Pachytesta to that of 
Lagenostoma has been recently pointed out by Professor Oliver 1 , and 
a transverse section has been constructed which exhibits its compartmental 
nature at a level much lower than that in which it can be demonstrated 
in Lagenostoma. Professor Oliver adds : { The presence of vascular strands 
in the chambers of Lagenostoma is the most important difference.’ 
Much fuller details are to hand of another seed which seems to bear 
out this theory. I refer to Bennettites Morierei , Sap. and Mar. (spec.), 
which has been admirably worked out by Professor Lignier 2 . This 
fructification, as is well known, belongs to a much later horizon, i. e. 
Mesozoic, and shows Cycadean affinities. 
If one consults Lignier’s Plate III, Figs. 35 and 37, one sees transverse 
sections of the upper part of the seed, showing the thick integument 
divided up into four compartments by radiating vertical walls of flattened 
cells, very comparable to those which separate the constituent members of 
a synangium. 
The interior of each compartment is described as succulent tissue, 
but offers an abrupt contrast to the walls. Plate III, Fig. 38, shows 
1 Oliver, ‘ On some Points of apparent Resemblance in certain Fossil and recent Gymnospermous 
Seeds.’ New Phytologist, vol. i, p. 150, Text-figure 5. 
2 O. Lignier, ‘Structure et Affinit^s du Bennettites Morierei , Sap. and Mar. (sp.).’ VegCtaux 
fossiles de Normandie. Caen, 1894. 
