43 8 Lawson . — The G a met op hy tes , Fertilization and 
shown in Fig. 59 , the tips of the suspensors usually separated from each 
other. There were generally one or two embryo-cells at the end of each 
suspensor. The two cells at the tip were merely the two-celled stage of 
the embryo proper. In no case was I able to find more than one embryo 
developed from a single suspensor as Coker (’03) has recently reported for 
Taxodium. One embryo was generally developed from one suspensor, but 
very frequently a single embryo was found at the united tips of two or 
sometimes three suspensors. 
In the later stages, distinct embryonal tubes are developed from the 
cells nearest the suspensors, in much the same fashion as they do in 
Taxodium (Coker, ’03). They are, however, not as numerous or as long 
as in Taxodium . 
While differing in certain minor details, the development of the 
embryo of Cryptomeria and Taxodium is very similar. To Sequoia , where 
there are no free nuclei formed in the pro-embryo, Cryptomeria bears no 
resemblance whatever. 
Summary. 
The reduction-division which leads to the formation of the tetrads 
takes place during the latter part of October, although pollination does 
not take place until March of the following spring. At the time of 
pollination the microspore contains a tube-cell and a generative cell. Cells 
or nuclei representing the vegetative tissue of the male gametophyte are 
not formed. 
There are usually four or five microspores deposited on the nucellus at 
the base of the micropyle. These all germinate and produce pollen-tubes, 
which penetrate the nucellar tissue at the apex. At the time of penetration 
the generative nucleus divides, so that the young pollen-tube contains the 
tube-, stalk-, and body-nuclei. 
The body-nucleus very soon enlarges and becomes surrounded by 
a dense zone of cytoplasm and starch-grains. A membrane is formed 
at the periphery of this zone. The pollen-tube now contains one large cell 
and two free nuclei. 
After the tip of the pollen-tube has reached the depression above the 
archegonium-complex, the body-cell divides and gives rise to two male cells. 
The two male cells when mature are spherical in form, of equal size, and 
both are functional. They enter separate archegonia. 
There are three or four macrospore-mother-cells differentiated in the 
nucellus at a point just a little above the insertion of the integument. Each 
mother-cell divides twice, and there are consequently from twelve to sixteen 
macrospores formed. 
Only one of the macrospores germinates and develops into the female 
