in Blechnum and Osmunda. 
47 
is perfectly enormous and much greater in quantity than in 
Blechnum . In Osmunda no fungus was found to accompany 
the young leaves, the increased amount of tannin being perhaps 
unfavourable to the mycelial growth. 
The fate of the woolly hairs is quite different from that of 
the glands, notwithstanding the similarity in the mode of 
development. Their cell-wall, which in the adult cells is often 
of very appreciable thickness, permanently retains its cellulose 
character and does not become cuticularised. The starch- 
grains gradually disappear from the mature cells, and on some 
of the transverse walls a callus may occur. In many of the 
cells the contents especially devote themselves to the forma- 
tion of tannin, in which case they appear clear and highly 
refractive. In the very oldest cells the contents entirely 
disappear. With these remarks we end our description of 
Osmunda. 
General Results. 
The secretory hairs which we have now described are 
characterised by a peculiarity rarely met with in glandular tri- 
chomes, in that the secretion does not leave the cell but remains 
included in the protoplasmic body throughout its entire life. 
The continuous aggregation, the manner of secretion, and the 
dense and solid character of the substance thus formed, enable 
us to understand how it is that in these particular glands the 
structure is clear and well-defined. 
The secretion commences by the breaking down of a 
portion of the innermost layers of the endoplasm at a number 
of contiguous but isolated areas. The first secretion occurs 
just beneath the free surface, and takes place equally around 
the whole cell-cavity, and the phenomenon steadily continues 
from within outwards, extending to deeper and deeper layers, 
until the whole of the endoplasm, together with the substance 
of the plastids, have taken their share in the process. We 
are unable to state whether these changes are limited to the 
endoplasm only, since the exact boundary between the ecto- 
plasm and endoplasm cannot be defined. In the present state 
