2j6 Bower. — Some Normal and Abnormal 
be seen that the main features are the same in both species, 
especially with respect to the external appearance suggesting 
a number of segments cut off from an initial cell. Again, 
in the antheridia of Hymenophyllum Smithii described and 
figured by Goebel 1 , an alternation of the basal walls is indi- 
cated, not unlike that which appears to obtain in Tr. pyxidi- 
ferum. 
In the case of the antheridia above described, we should be 
justified in regarding the whole antheridium with its pedicel 
as replacing a lateral branch, the terminal cell of which is 
represented by the head of the antheridium. 
The archegonia^ on the other hand, are borne on massive 
growths (archegoniophores), which may be held in some cases 
to represent the apical cell of a lateral branch (Fig. io), or the 
whole of a lateral branch (Figs. 11,12). In one case however, 
which has a special interest, the mass of tissue on which the 
archegonia are produced is obviously the result of partition of 
a single cell of a filament, without any marked increase of size 
having taken place (Fig. 13). This may be regarded as the 
simplest form of an archegoniophore. hitherto described in any 
Fern, or even in any Bryophyte, and it approaches near to 
that suggested by Goebel 2 as the simplest possible, in which 
the sexual organs would be inserted directly on the protonemal 
threads. Thus, while there is an entire absence of flattened 
developments on which the archegonia might be borne, as 
described for some other allied forms 3 , the actual position and 
origin of the archegoniophore is subject to some variation. 
Further, the cell-divisions (as far as I have been able to trace 
them) which give rise to these massive growths follow no 
definite order ; this may be gathered from a comparison of the 
specimens represented in Figs. 11, 12, 13, 14. A wedge-shaped 
cell is sometimes to be seen at the apex of the growth, but, 
even if this be a true initial cell, its activity is not long 
continued, nor is it always to be found (Fig. 13). 
1 1 . c. Taf. xiii. Fig. 89. Compare also diagram, Taf. xiii. Fig. 82. 
2 1. c. p. 94. 
3 Cf. Mettenius, 1 . c., Tr. sinuosum, etc. 
