The Comparative Anatomy and Phylogeny of the 
Cyperaceae 
BY 
AMON B. PLOWMAN. 
With Plates I and II and two Figures. 
Synopsis. 
PAGE 
1. Introduction 2 
2. Problems 2 
3. The Root 3 
4. The Rhizome 4 
a. General Morphology ............ 5 
b. The Amphivasae ............. 6 
c. The Centrivasae 7 
d. Generic Key .............. 9 
e. Descriptive Table 12 
5. The Aerial Stem 10 
a. General Morphology ............ 11 
b. Mode of Origin and Apical Growth . . . . . . . . .11 
c. Internal Anatomy ............ .11 
aa. The Chlorocyperaceae 1 4 
bb. The Sclerocyperaceae . . . 15 
cc. Occurrence of Cambium . . . . . . . . . .18 
d. Descriptive Table .16 
e. Nodal Complexes ..... 18 
f Course of Fibro-vascular Bundles 18 
g. Cortical Bundles 2 3 
h. The Scapose Type 21 
i. Axillary Buds ........21 
6. The Leaf 21 
a. General Morphology ............ 21 
b. Internal Anatomy ... .......... 22 
c. Mode of Insertion ............. 2 2 
d. Dominance in Stelar Development . . . . 23 
7. The Floral Axis 23 
a. Morphology and Anatomy . . . . . . . . . . . 23 
b. Dicotyledonous Arrangement of Bundles 23 
8. Fossil Cyperaceae 24 
9. Phylogenetic Considerations 24 
a. Inter-relationships of the Cyperaceae . . . . . . . . . 25 
b. Position among the Monocotyledons ......... 26 
c. Bearing upon the Phylogeny of the Monocotyledons 26 
10. Summary and Conclusions 27 
11. Literature 28 
12. Explanation of the Plates 30 
1 Contributions from the Phanerogamic Laboratories of Harvard University, No. 2. 
[Annals of Botany, Vol. XX. No. LXXVII. January, 1906.] 
B 
