Some Points in the Morphology of Phyllocladus 
alpinus, Hook. 
BY 
AGNES ROBERTSON, D.Sc. 
Quain Student in Botany , University College , London. 
With Plates XVII and XVIII. 
I. Introduction. 
T)HYLLOCLADUS is a peculiar and isolated genus belonging to the 
^ Taxaceae, and characterized by the reduction of its true leaves 
to pointed scales, and the expansion of certain of its stem-branches into 
flattened leaf-like structures. Pilger in his monograph of the Taxaceae 1 
places the genus in a sub-family by itself, intermediate between Podo- 
carpoideae and Taxoideae. There are six species confined to New Zealand, 
Tasmania, and the Indian Archipelago. The isolated systematic position 
of the genus, its restricted distribution and curious external morphology, 
added to the fact that little was known of its anatomy or development, 
suggested to Prof. F. W. Oliver that it might prove an interesting subject 
for investigation. Accordingly, through the kindness of Dr. L. Cockayne, 
who had access to Phyllocladus alpinus , the Celery Pine of New Zealand 
in cultivation in the Christchurch Botanic Garden, five gatherings of 
material were sent to University College during the years 190a, 1903, and 
1904, and were handed to me for examination. The following account 
is based upon this material, supplemented by a supply of the vegetative 
organs of two other species, P. rhomhoidalis and P. trichomanoides , which 
Mr. R. I. Lynch, M.A., was so kind as to obtain for me from the Edinburgh 
Botanic Garden, and which has been very useful for comparison. The 
material of P. alpinus , which was fixed in methylated spirit, was unfor- 
tunately inadequate for anything like a complete investigation, and it 
is greatly to be desired that some one who has access to the genus in 
its native haunts should give us an exhaustive account of its mor- 
phology. I am much indebted to Prof. Oliver for his kind help and advice 
throughout the work. 
1 R. Pilger, Das Pflanzenreich, iv, 5, Taxaceae, 1903. 
[Annals of Botany, Vol. XX. No. LXXIX. July, 1906.] 
