The Wound Reactions of Brachyphyllum l . 
BY 
EDWARD C. JEFFREY. 
Assistant Professor of Botany in Harvard University. 
With Plates XXVII and XXVIII. 
‘HE genus Brachyphyllum was established by Brongniart for the leafy 
A branches of a Jurassic coniferous plant of doubtful affinities (Prod. 
Hist Veg. Fossiles, p. 109, 1828). Saporta, who subsequently described 
a number of species of the genus (Plantes Jurassiques, vol. iii, 1884), placed 
it with the Araucarineae. This type of foliar shoot from the Cretaceous 
and Jurassic deposits of different parts of the world has been referred 
by various authors under several generic appellations to the Araucarineae, 
Cupressineae, and Sequoiineae, with perhaps the weight of authority in- 
clining towards the last attribution. The present writer, in collaboration 
with Dr. Arthur Hollick of the New York Botanic Garden, on the basis 
of well preserved material, has been able to settle finally the systematic 
position of these interesting and omnipresent relics from the Jurassic and 
Cretaceous strata, from the investigation of internal structure. The anato- 
mical features of Brachyphyllum , as exemplified by the North American 
species B. macrocarpum , N ewberry, are such that there can now be no doubt 
as to its Araucarian affinities (Affinities of certain Cretaceous Plant remains 
commonly referred to the genera Dammara and Brachyphyllum, American 
Naturalist, vol. xl, pp. 189-215, March, 1906). In the article just cited, 
it has been demonstrated that Brachyphyllum , although presenting un- 
doubted Araucarian features, differed markedly from existing and many 
extinct Araucarineae, both in the structure and wound reactions of its 
wood. These features are to be farther discussed in the present article, 
because they appear to be of considerable importance not only from the 
standpoint of the identification of lignites and mineralized woods, which 
are the remains of Brachyphyllum , but also from that of the general ques- 
tion of the affinities and phylogenetic history of the Araucarineae. The 
present writer has now in his possession a considerable amount of fossil 
1 Contributions from the Phanerogamic Laboratories of Harvard University, No. 7. 
[Annals of Botany, VoJ. XX. No. LXXX. October, 1906.] 
