1 86 Atkinson . — A Monograph of the 
ends of the ray-cells and extend both up and down to the 
bases of the papillae. Fig. u is from Lemanea (S acker ia) 
fucina. Fig. 32 represents a part of a longitudinal section of 
the sexual shoot of a species of the subgenus Lemanea ( L . 
australis Atk.) : a is the central axis, b U b" b rer the ray- 
cells ; d and d' a descending and ascending generative fila- 
ment. The descending one ends, as can be seen in the figure, 
at the base of the antheridia g. In Figs. 32- and 11 e e e are 
the female organs, procarps. In Fig. 32 they arise from the 
generative filaments near the middle of the interval between 
two adjacent antherid-bands, and in Fig. 11 they arise from 
the generative filaments near the antherid-papillae. 
In order to be concise in speaking of the different portions 
of the sexual shoot and save the circumlocution which I have 
used in speaking of the ‘ interval ’ and ‘ whorls of papillae, 5 I 
will use definite terms for the alternating zones. 
Since the terms ‘ node 5 and ‘ internode ’ do not in the 
usual acceptance of those terms seem proper, and because I 
wish to use terms which have some appropriate significance 
of the function of the alternating zones, I shall use the term 
antherid-zone for that portion of the sexual shoot which 
includes the antherid-band or antherid-papillae as the case 
may be ; and the term procarp-zone for the alternating zones, 
or that portion which lies between two adjacent antherid- 
zones. Then since a procarp-zone with half of each adjacent 
antherid-zone is a sexually complete segment of the sexual 
shoot, I shall use the term sexual segment, or segment, to in- 
dicate that. 
While in general the structure of the apparatus of fructifi- 
cation is similar in both subgenera, its elaboration in the sub- 
genus Lemanea is more complex, and each merits considera- 
tion in detail. In the subgenus S acker ia the central axis is 
always naked. The ray-cells are T-shaped or L-shaped, one 
end of the cell being joined to the central axis, the two other 
ends forming the beginnings respectively of the ascending and 
descending generative filaments and closely applied to the 
medullary layer. Each ray-cell is ‘ tied 5 to the medullary 
