Oliver . — On Sar codes sanguined, Torr. 321 
made use of by the allied genera. At the time of flowering, 
there is around the base of the style a ring-like depression, or 
invagination, of the epidermis. This is seen in the longitudinal 
sections of the ovary represented in Figs. 1 7 and 1 8 ; also in 
the transverse section, Fig. 21, C , taken at the insertion of the 
style. In a more highly magnified longitudinal section (Fig. 
19) the nature of the depression is seen. It penetrates three 
quarters of the way through the ovary wall, and only a narrow 
zone of tissue separates it from the cavity of the ovary. It is 
the small cells ( 'x , Fig. 19) of this region that, breaking down 
as the fruit ripens, bring about the peculiar dehiscence (circum- 
scissile) described above. In Monotropa a circular depression 
occurs in the same position, but its depth is only one-third of 
that in Sarcodes , and it does not lend itself to this method of 
dehiscence. Similarly, in some other genera of Monotropeae. 
In old specimens of Sarcodes , when many of the fruits are 
already in seed, small flowers appear in the axils of the fleshy 
leaves, below the general inflorescence. These, in ordinary 
flowering specimens as that figured, are still minute buds, 
1 mm. in length, embedded in the fleshy bases of the scale. 
Development of the Ovules . — The ovules are very nu- 
merous, and their development can be easily followed. 
Each arises on the young placenta as a small upgrowth of 
tissue, by the formation of tangential divisions in the layer 
immediately below the epidermis. At an early period the 
terminal cell of the hypodermal tissue of the young ovule is 
marked out by its relatively large size (Figs. 24 and 25, a), 
and as the terminal portion of the ovule (nucellus) begins to 
curve over, it becomes very prominent. At the same time, 
the commencement of the single coat of the ovule arises (Fig. 
25, int). The large terminal cell is the archesporium , and it 
undergoes a number of divisions, prior to the formation of the 
embryo-sac. These are shown in Figs. 26-30. There is, cut 
off from the apical end of the archesporium, first one and then 
a second £ cap-cell 5 ( cc in Figs,) ; this number is not exceeded. 
The large, third cell of the row, so formed, is the embryo-sac. 
In succeeding stages the cap-cells gradually dwindle as the 
