60 
PACIFIC SCIENCE, Vol. V, January, 1951 
1. Myoporum sandwicense A. Gray, Proc. 
Am. Acad. Sci. 6: 52-53. 1866. 
Polycoelium sandwicense A. DC., Prodr. 11: 
706. 1847 (illegitimate name). 
Usually much-branched shrubs 1-3 m. 
high, but stature varying from prostrate 
(littoral form) to trees 15 m. tall. Stem usually 
glabrous with smooth green or olive-brown 
bark when young, gray and cracking into 
chunks when older. Leaves alternate, often 
lanceolate but varying from linear-lanceolate 
to ovate, acute to long-acuminate, entire or 
often crenate or serrate, decurrent-petiolate, 
3.5- 22 cm. long, 0.5-4 cm. wide; usually 
somewhat fleshy but varying from submem- 
branous to stiff-coriaceous when dry; usually 
glabrous or more rarely pubescent. Flowers 
axillary, in clusters of 2-10, often 1 or more 
abortive; pedicels terete or flattened, 0.5- 
1.7 cm. long. Calyx lobes 4-9, mostly 5, 
1- 6 mm. long, broadly ovate to oblong 
or lanceolate, green and herbaceous or 
slightly scarious on the minutely glandular- 
denticulate margins, glandular-punctate, usu- 
ally glabrous. Corolla 4-9- (mostly 5-) lobed, 
4.5- 12 mm. long, campanulate to funnelform, 
usually glandular-punctate, entirely glabrous 
to densely pubescent within; color pure 
white, white with pink-purplish splotches, 
or entirely pink; corolla lobes ovate- deltoid 
to oblong, emarginate, obtuse, or apiculate, 
2- 7 mm. long. Stamens inserted on the 
corolla, the same number as and alternate 
with the corolla lobes or occasionally 1 or 2 
fewer; filaments flattened-subulate, 1.5-5 
mm. long; anthers 0.5-1. 5 mm. long, the 
cells confluent by a common suture and thus 
changing from parallel to widely divergent 
at an thesis. Ovary cylindric-conic to de- 
pressed-conic, 1.5-4 mm. long, glabrous, 
green, 4-12-celled, the base with a yellow or 
purple apparently nectariferous ring; style 
terete or flattened, 1.5-6 mm. long; stigma 
convex, obscurely 2-5-grooved. Drupe fleshy, 
the exocarp greenish-white or creamy-white, 
rarely pinkish or purplish; endocarp 2-10 
mm. long, 2-9 mm. broad, smooth or ribbed, 
subglobose or variously flattened, hard and 
bony when mature. Seeds one in each cell, 
spindle-shaped, papery when dry, 2-2.5 mm. 
long. 
The name Myoporum sandwicense dates 
from its use by Asa Gray in 1866, the origi- 
nal combination Polycoelium sandwicense A. 
DC. being illegitimate under the International 
Rules of Botanical Nomenclature, Article 
60 (1). The generic name Polycoelium was 
a substitution for the earlier Pentacoelium of 
Siebold and Zuccarini, and hence was super- 
fluous when published. According to Article 
69 of the rules, Gray’s adoption of the epithet 
sandwicense is treated as a new name and not 
as a new combination. 
As here construed M. sandwicense em- 
braces three rather diverse subspecies, each 
of which could well be treated as a species if 
their differences were constant. I have been 
reluctant to reduce M. Wilderi to a subspecies, 
but if this were not done consistency would 
demand the elevation of ssp. St.-Johnii to 
specific rank, and this action does not seem 
warranted. 
2. Myoporum sandwicense ssp. sand- 
wicense. 
Myoporum sandwicense A. Gray, Proc. Am. 
Acad. Sci. 6: 52-53. 1866 (in part, excluding 
reference to Douglas collection). 
Prostrate shrubs to tall trees. Leaves mostly 
lanceolate, less commonly almost linear or 
ovate or elliptic, never spatulate, 3.5-22 cm. 
long, 0.5-4 cm. broad, glabrous or pubescent. 
Flowers in clusters of 2-10 (mostly 3-5). 
Calyx mostly 5 -lobed, lobes broadly ovate to 
oblong or lanceolate, 1-5 mm. long. Corolla 
mostly 5 -lobed, glandular-punctate, glab- 
rous or pubescent within, 4.5-12 mm. long. 
Stamens isomerous with the corolla lobes, the 
fifth one occasionally reduced or absent. 
Ovary 4-10-celled, 1.5-4 mm. long; style 
glabrous, 1.5-6 mm. long. Drupe creamy- or 
greenish-white or rarely pinkish or purplish; 
