134 
PACIFIC SCIENCE, VoL V, April, 1951 
paratypes (10 males and 8 females): Atherton 
Tableland, Queensland, Nov., 1949, ex 
Aglaia ferruginea. Also, two paratype females: 
near Atherton, Queensland, Feb., 1950, ex 
Aglaia sapindina. 
Holotype, allotype, and four paratypes 
have been deposited in the United States 
National Museum. The remainder of the 
paratypes are being distributed to the fol- 
lowing museums and individual collections: 
Australian Museum, Sydney, Australia; Bishop 
Museum, Flonolulu, T. H.; Board of Agri- 
culture and Forestry, Honolulu, T. H.; 
Hawaiian Sugar Planters’ Association, Hono- 
lulu, T. H.; University of Hawaii, Honolulu, 
T. H.; the collection of Dr. H. K. Munro, 
Pretoria, South Africa; and the collection of 
Dr. M. Hering, University of Berlin. 
Dacus (Heterodaculus) n. subgen. 
This subgenus is very closely related to 
Dacus {Daculus) but is differentiated by the 
lack of the supernumerary lobes in the wings 
of the males. If the generic concepts used 
by various other workers in this group 
were followed, this difference would justify 
setting up a new genus for the species at 
hand. I am not in favor of this procedure and 
prefer that genera not be based upon second- 
ary sexual characters unless accompanied by 
ample supporting characters, preferably char- 
acters common to both sexes. The wing 
venation is considerably different from that of 
known species of the subgenus Daculus, and, 
on the basis of a comparison of Dacus 
{Heterodaculus) vtsendus n. sp. with the de- 
scribed species of the subgenus Daculus, it 
would appear that they should be placed in 
different genera. However, on the basis of a 
broad comparative study of the characters 
throughout the subfamily Dacinae, these 
characters do not appear to be of more than 
subgeneric importance. The variations and 
intergradations of the intrageneric characters 
of the genus Dacus sens. lat. encompass the 
structural differences seen in the species at 
hand. 
Aside from the secondary sexual characters 
mentioned above, the subgenus differs from 
Daculus and other closely related subgenera 
by having the r-m crossvein situated below 
the middle of the discal cell. The section of 
cell R from the fumose portion (directly 
above the base of vein M3+4) to the r-m cross- 
vein is very short compared to the same sec- 
tion in Daculus and most other Dacus. This 
portion of cell R is slightly shorter than cell 
M (Fig. 8c). In all the known species of 
Daculus and almost all known species of 
Dacus the r-m crossvein is situated beyond the 
middle of the discal cell, and the apical por- 
tion of cell R (beyond the fumosity) is 
about one and one-half times longer than cell 
M (Fig. 5^). In Heterodaculus the apex of vein 
Ri-I -2 is well beyond a point opposite the r-m 
cross vein; the top of the crossvein is almost 
opposite the middle of the third section of 
the costa. In Daculus and other Dacus, with a 
few exceptions, the top of the r-m crossvein 
is about opposite the apex of vein R 1+2 and 
the bottom is slightly beyond its apex. At 
least three species of Dacus {Strumeta) — re- 
currens (Hering) , pulcher (Tryon) , and manskii 
(Perkins and May) — have the r-m crossvein 
located in this more anterior position, so this 
would probably not be a valid generic char- 
acter. The apical portion of vein M3-1-4 is more 
elongate and the lower corner of cell 1st M 2 
is farther from the wing margin than in other 
Dacus species which I have studied. The vein 
reaches four-fifths of the distance to the wing 
margin, and the distance from the lower cor- 
ner of cell 1st M 2 to the margin is equal to 
about one- third the length of the m crossvein. 
The head possesses three to four pairs of 
inferior fronto-orbital bristles. A pair of small 
but conspicuous postocellar bristles is also 
present. 
GENOTYPE: Dacus {Heterodaculus) vtsendus 
n. sp. 
