170 
Cairns, same host and date as above, mango, 
Dec., 1949, and guava, Apr., 1950; across 
inlet from Cairns, ex ’'fruit plant No. 138,”® 
Feb., 195 0; and near Hartley’s Creek, N. 
Queensland, ex Clausena brevistylus, Feb., 
1950. note: In the longer series of specimens 
from the above localities and hosts the humeri 
vary in color from brown to yellow. 
Dacus tryoni var. sarcocephali (Tryon) 
Chaetodacus tryoni var. sarcocephali Tryon 1927. 
Roy. Soc. Queensland, Proc. 38(14): 188. 
As discussed above, this is just a color 
variation of D. tryoni, and in many of the 
series at hand so many borderline specimens 
are present (also verging into var. melas) that 
it does not seem practical to break them 
down even into varieties. Since vatiety 
sarcocephali apparently is commonly accepted 
by the Australian workers, I am including it in 
this discussion. The variety is supposed to 
differ from typical tryoni by having distinct 
black markings on the mesonotum, accord- 
ing to Perkins and May, "comprising two 
dots and two streaks.” They also say that 
the flies are mostly smaller in size. The size 
difference apparently is of no importance 
as no constant difference in size has been ob- 
served in the long series at hand. The oviposi- 
tors are identical with the typical form. Mr. 
May offered the following comments in one 
of his letters, "Re Tryon’s 1927 paper and 
the varieties of S. tryoni: I have yet to go into 
this problem. . . . Should you regard var. 
rrnisa and vda. juglandis as true synonyms, then 
you would also treat var. sarcocephali on the 
same basis? I have yet to find differences of a 
structural nature, between tryoni and tryoni 
var. sarcocephali. Var. jiiglandis is quite a small 
fly in comparison to tryoni but that could be 
due to food.” I may not be correct in treating 
the forms in this tryoni complex as I have, 
®Mr. Krauss reported that this plant was determined 
(from fruit only) as Terminalia melanocarpa, but he was 
doubtful of this determination. He stated that it came 
from a "large tree . . . dark purple fruit, Vi inch diameter, 
M inch long, juicy purple pulp.” 
PACIFIC SCIENCE, Vol. V, April, 1951 
but the procedure followed here seems to be 
the most logical way of handling them. 
TYPE LOCALITY: Brisbane, ex Sarcocephalus 
cordatus and Psidium sp. 
Type in the Queensland Museum. 
Typical sarcocephali are in the collection 
from Hartley’s Creek, N. Queensland, ex 
Polyalthia nitidissima and Ganophyllum falca- 
tum, Feb.-Mar., 1950 (some of these inter- 
grade into var. melas)\ Atherton Tableland, 
Queensland, ex Khodamnia sessiliflora and 
Solanum seaforthianum, Mar., 1950; Cairns, 
Queensland, ex guava. Mar., 1950 (inter- 
grading into var. melas') ; near Hartley’s Creek, 
N. Queensland, ex Terminalia melanocarpa. 
Mar., 1950 (smaller specimens intergrading 
in color toward var. melas)’, Ellis Beach near 
Cairns, Queensland, ex "plant 129,’”^ Feb., 
I95O; also mixed in with a large share of 
the typical tryoni reported above. 
ACANTHONEURA Macquatt 
Acanthoneura Macquart 1843. Dipt. Exot. 
3(3): 220. 
This genus is closely related to Pioxa 
Walker but the aristae are short haired and 
just one pair of inferior fronto-orbital bristles 
is present. According to Malloch (1939) 
three species of Acanthoneura occur in 
Australia. Just one species, A. australina 
Hendel, was represented in the Krauss col- 
lection. It now appears that a fourth species, 
A. bicolor (Mq.), should be included here. 
GENOTYPE: Acanthoneura fuscipennis Mac- 
quart. 
KEY TO KNOWN AUSTRALIAN Acanthoneura^ 
1. Abdomen entirely black; thorax yellow- 
brown without black markings; 
wings with just one hyaline incision 
on the costa just beyond end of vein 
Ri +2 and with two hyaline spots in 
'^The plant "was apparently lost en route to Mr. 
C. J. White, Government Botanist, Brisbane.” Mr. 
Krauss said it came from a "small tree . . . small red 
fruit, ^6 in. long, Va in. diameter, in large clusters.” 
^Adapted from Malloch (1939<^). 
