Hawaiian Species of Scaevola — St. John 
not marcescent; calyx not bracteate 
at base CAMPHUSIA 
Corolla tube straight or nearly so and 
on upper side split to base; filaments 
early marcescent; calyx usually bi- 
bracteolate at base SCAEVOLA” 
Among the Hawaiian species of Scaevola, 
the species 5. glabra is conspicuous by having 
a strongly curved, fleshy corolla tube which 
is cleft on the proximal side only about half- 
way down and the calyx or hypanthium or 
base of the inferior ovary is naked or bractless 
at base. The question is, are these generic 
differences.^ 
The corolla is cleft about halfway, but it 
has a suture from there down to the base on 
which it may split in age. This seems an ex- 
treme specific character, but not a generic one. 
The calyx or ovary is bractless at the base, but 
so also are the numerous flowers in the large 
cyme of 5. frutescens (Miller) Krause of the 
Pacific and Indian Oceans, and so are they 
bractless in S. Helmstt E. Pritzel of Western 
Australia. The filaments of S. glabra are re- 
tained within the partly closed corolla tube 
and hence are little exposed. The filaments of 
most other Scaevola species are pushed outside 
as the slit tube of the corolla yields to the 
stress and turns to the anterior side and more 
or less flattens out, so the exposed filaments 
after anthesis wither and dry. However, no 
important difference between marcescent and 
non-marcescent filaments has been noted. 
The yellow color of the corolla is not unique, 
for other species have yellow flowers, as, for 
example, S. enantophylla F. von Mueller. Other 
species have their corollas of varying textures 
from membranous to fleshy, the Hawaiian 
species S. coriacea Nuttall having the whitish 
flowers of the 1- to 3-flowered cymes definite- 
ly fleshy. Other species, such as S. tomentosa 
Gaudichaud of Western Australia, have 
curved corollas. In some families the curving 
of the corolla provides good characters, but 
in Scaevola, with various species having the 
corolla tubes straight, nearly straight, or 
slightly or markedly curved, the attempt to 
31 
set off one species as a segregate genus by the 
use of this character is patently unsound. The 
key and diagnostic characters used by 
Degener for Camphusia are judged equally in- 
valid as the one of De Vriese. In conclusion, 
Camphusia seems correctly placed as a syno- 
nym of Scaevola, as it was by nearly all 
botanists and as it was in the monograph by 
Krause (1912: 119, 130) and by Skottsberg in 
his earlier writing (1927: 39), though in a 
later paper (1939: 184) there is an indication 
that he intends to restore Camphusia. 
No specimens of any plant like S. glabra 
from the island of Maui are available. Krause 
lists one collected by Hillebrand and now in 
the herbarium at Wien. This has not been 
available for study, and, like Skottsberg and 
Degener, the writer has seen no recent collec- 
tions from that area. If it really came from 
Maui, it might well be a distinct kind. 
Since S. kauaiensis is here accepted as a 
species, it is desirable to print a description 
of it: 
Shrub up to 4 m. tall; branches forking but 
often elongate and wand-like, the bark 
smooth, pale brown with raised corky lunate 
leaf scars and branch scars; branchlets smooth 
leafy, often for a considerable distance, stout, 
1-15 mm. in diameter; axils with a prominent 
hairy tuft, the abundant hairs tawny, silky 
lanate, 6-7 mm. long, tardily deciduous after 
the fall of the leaves; leaves glabrous, alter- 
nate, or in part subopposite; petioles 12-40 
mm, long; blades 5-12 cm. long, 15-55 mm. 
wide, coriaceous, oblanceolate to oval, the 
apex acute, the base cuneate or .decurrent, 
above dark green, below pale green; cymes 
axillary, reduced, and 1- or 2-flowered; pe- 
duncle 2-10 mm. long, glabrous except for 
the hairy tufts at the apex; bracts 2, opposite, 
terminal, 2-4 mm. long, lanceolate to ellip- 
tic, fleshy, with their axils tawny hairy tufted; 
pedicels 7-35 mm. long, glabrous, ascending 
or spreading, bractless; hypanthium 4-6 mm. 
long, narrowly obpyramidal, glabrous; calyx 
lobes 5-8 mm. long, linear or, rarely, lance- 
linear; corolla yellow, fleshy, curved; corolla 
